F. J. Meggitt 
197 
Familie der Ichthyotaenien, aufweist,” proceeding then to elaborate the differ¬ 
ences between the species of the same genus. The position of the female glands 
in the species of a genus is not constant. In the genus Bavainea Blanchard 
1891 they customarily lie on the median axis of the strobilus, but in D. lepto- 
trachela Hungerbiihler 1910 they are poral in position, nearly touching the 
longitudinal excretory vessels, and again in D. tetragona (Molin 1858) and D. 
echinobothrida (Megnin 1880) are distinctly poral. With regard to the size of 
the cirrus-sac, in his account of the type species, T. lemuris, Beddard (1911 a) 
gives neither measurements nor a statement of its extent, merely stating 
(1912, p. 607) “ Cirrus-sac rather large,” a vague statement unsuitable for use 
as a generic character. Neither do the three characters taken in combination 
assume sufficient importance to be of generic rank. From Beddard’s de¬ 
scription therefore, unless subsequently further distinctive characters be 
forthcoming, I would suggest the placing of T. lemuris in the genus Zschok- 
Jceella, —the egg-capsules being formed altogether independently of the uterus 
(1911, p. 1000)—and the suppression of the genus Thysanotaenia. 
For the identification of the two remaining genera Beddard (1912, p. 607) 
has provided a somewhat unsatisfactory key. He places I. hyracis (Ball.) 1 in 
the genus Zschokkeella and then distinguishes between the two genera by 
crediting Inermicapsifer with posterior genital pores, two groups of testes 
entirely separated from each other, a vas deferens together with a seminal 
network, and the absence of a well-developed uterus, and Zschokkeella with 
median genital pores, a continuous row of testes, the absence of a vesicula 
seminalis (and presumably of a seminal network?) and the presence of a well- 
developed uterus. With regard to the first character, the disposition of the 
testes, there is every variation between the type of I. setti v. Janicki 1910 
with the testes in two clearly separated groups, through Z. gambianum 
Beddard 1911 (1912, p. 582), where three testes form a connecting link be¬ 
tween the two groups to I. hyracis (Pall.) where they are arranged in a con¬ 
tinuous band along the posterior border of the segment, and I. paronae 
Bischoff 1912, where they are equally distributed throughout the whole pro¬ 
glottis, extending anteriorly to the genital ducts. For Zschokkeella muricola 
Baylis (1915, p. 47) states: “The testes are divisible into two groups; but they 
so nearly form a complete series across the segment (except where interrupted 
by the female organs) that it is difficult to say whether their arrangement is 
more like that seen in Inermicapsifer or that in Zschokkeella. In this respect 
the species seems to be intermediate between the two genera.’ A character 
so liable to variation as this cannot be considered adequate for the dis¬ 
crimination of genera. The further distinction drawn by means of the vesicula 
seminalis is inaccurate. Fuhrmann (1902, p. 139) for Z. linstowi, the type species 
1 Taenia hyracis Pallas 17G7 was selected by von Janicki (1910) for the type species of his 
genus Inermicapsifer. If this species be removed to the genus Zschokkeella (Fuhrmann 1901), 
the name Inermicapsifer falls into synonymy with Zschokkeella and Beddard s first group, with 
testes in two distinct groups, should be renamed. 
