198 Cricetomys gambianum 
where, according to Beddard’s table, it should be absent, states: “Am inner 
Ende der schwach muskulosen Penistasche liegt eine kleinere Vesicula semi- 
nalis.” On the other hand, Baylis (1915, p. 43) found it absent in Z. muricola 
while, with the exception of 7. capensis Beddard 1912 and the new species 
just described, it is absent or, being unmentioned, presumably absent in all 
the nominal species of Inermicapsifer, whether with a continuous band of 
testes and therefore referable to Zschokkeella, or with two groups and therefore 
referable to Inermicapsifer. The position of the genital pore, the third cha¬ 
racter, is anterior in I. parvulus Bischoff 1912, and posterior in 7. setti v. 
Janicki 1910—both with two separate groups of testes—and anterior in Z. 
remota v. Linstow 1906 and posterior in 7. parona Bischoff 1912—both with 
a continuous band of testes. The last character in the key, uterus well or little 
developed, is a comparative one and therefore suffers from vagueness. In 
7. hyracis the uterus is stated Janicki (1910, p. 381) to extend between the 
excretory vessels as a broad sack with outgrowths, disappearing to form the 
walls of the primary egg-capsules: in 7. capensis the oviduct is stated (Beddard 
1912, 586), to be “quite short and ends more or less abruptly in a strand of 
condensed parenchymal tissue.” In Z. gambianum (Beddard 1911, p. 659) the 
uterus is “not much more than a transversely running tube extending nearly 
right across the proglottis in which the eggs occur, but with which the uterus 
never appears to be stuffed. I could find no outgrowths of this centrally- 
placed uterus, and there was certainly nothing in the nature of a reticular 
formation of its cavity.” Other investigators are content merely to state, 
“Der Uterus, der sich auf jiingerem Stadium unregelmassig zwischen die 
Hoden einscheibt, lost sich etwa im letzten Yiertel der Strobila sehr rasch in 
Eikapseln auf ” (Bischoff 1913, p. 238): “The uterus persists through about 28 
segments, but after a short time appears to break down altogether, and the 
ova are seen to be scattered among the parenchyma ” (Baylis 1915, p. 44): and 
“Der Uterus bildet angangs ein median verlaufendes, leicht gewelltes Rohr, 
das beiderseits bis an die Langsnerven reicht. Seine Wandung verscheint bald 
und werden die Eier ins Parenchyme gestossen” (Fuhrmann 1902, 140). 7. 
capensis is the only species which can thus definitely be placed in the genus 
Inermicapsifer, the remaining 17 species falling into the genus Zschokkeella 
which then becomes a rather heterogeneous assemblage, for the testes may be 
continuous (7. hyracis) or in two groups (7. setti), a vesicula seminalis may be 
present (Z. linstowi) or absent (Z. muricola), and the genital pore posterior 
(7. paronae), or median (7. interpositus v. Janicki 1910), or anterior (Z. muri¬ 
cola) and there may be almost any permutation and combination of these 
characters. From the above facts it may be seen that it is impossible to use 
Beddard’s definition of the two genera unless it is prepared to reserve the genus 
Inermicapsifer for 7. capensis alone and make Zschokkeella a dumping ground 
for all the other species. 
Yon Janicki (1910, p. 394) distinguishes his genus from Zschokkeella by the 
following points: eggs in parenchymatous capsules and not merely in “Binde- 
