M. Koidzumt 
23 9 
termes lacteus from Australia, Schedorhinolermes putorius from Australia, and 
Schedorhinotermes intermedins from French Guinea. He referred the protozoa 
to 13 genera, 18 species, and 5 varieties—5 of the 13 genera, viz. Joenina , 
Slaurojoenia, Spirotrichonymphella , Pseudotrypanosoma , and Diplomympha , 
being newly established ones. 
In a work on the structure and biology of Archotermopsis of India, pub¬ 
lished in 1919, Imms gave descriptions of the intestinal protozoa harboured 
by that termite. Further studies of these organisms are being carried on by 
Cutler, who obtained a stock of the living termites from Imms, and two 
papers (1919 and 1920) have already been published, embodying his studies 
on species of Trichomonas (Ditrichomonas ) and Joenopsis (gen. nov.). Kofoid 
and Swezy published in 1919 a series of papers dealing with the structure 
and mode of multiplication of four species of intestinal protozoa harboured 
by Termopsis angusticollis of California. Each species was referred to a 
different genus, viz. Streblomastix, Trichomitus, Trichonympha and Leidyopsis , 
the first and the last being new. 
The physiological relationship between the termites and their intestinal 
protozoa is a highly important and deeply interesting subject. Do the 
organisms live in the intestine parasitically, commensally, or symbioticallv, 
as different authors have supposed? A study of this question has been made 
by Buscalioni and Comes (1910). They examined the reactions to chemical 
reagents of each region of the body of the protozoa, to determine the fate 
of the ingested wood fibres, and reported their results in detail. Imms (1919) 
has also devoted one chapter of his work to the consideration of some physio¬ 
logical questions. 
A note by Buttel-Beepen (1914) recording the results of his examination 
of many genera of termites in Java, Sumatra, Malacca, and Ceylon, to de- 
teimme the piesence oi absence of intestinal protozoa, will be suggestive for 
further studies; but his work, though adding to our knowledge of the dis¬ 
tribution of these protozoa, contains nothing else of interest for the present 
paper. 
A list of genera hitherto described will be given here. The genera estab¬ 
lished for the organisms harboured exclusively by termites, the type species 
of these genera, and their synonyms, are as follows: 
Genus. 
Trichonympha Leidy, 1877. 
Pyrsonympha Leidy, 1877. 
Dinenymj)ha Leidy, 1877. 
Joenia Grassi, 1885. 
Leidyonella Frcnzel, IS91. 
Micro joenia Grassi, 1892. 
Holomastigotes Grassi, 1892. 
Devescovina Foa, 1-905. 
Calonympha Foa, -1905. 
Gymnonympha Dobell, 1910. 
Type Species. 
T. agilis Leidy. 
P. vertens Leidy. 
D. gracilis Leidy. 
J. annectens Grassi. 
L. cordubensis Frenzel. 
M. hexamitoides Grassi. 
11. elongatum Grassi. 
D. striata Foa. 
C. grassii Foa. 
Gr. zeylanica Dobell. 
