M. Koidzumi 
255 
II. Pseudotrichonympha. 
Hartmann (1910) found three trichonymphids in a species of Coplolermes 1 
in Brazil, and interpreted them as representing the young, the female, and 
the male, of a single new species which he called "Trichonympha hertwigiP 
Soon after the appearance of Hartmann’s work, objections to his view were 
raised by Grassi (1911), and the “sexual forms” of Hartmann were made 
independent species, each being referred to a new genus, viz. Holomastigotoides 
and Pseudotrichonympha, proposed for the “female” and the ‘'male” forms 
respectively. (“Male” and “female” contrariwise misprinted at that time, 
and corrected later.) In an article contributed to the Handbuch der Natur- 
wissenschaften (Vol. hi. 1913), Hartmann subsequently adopted the generic 
name Holomastigotoides, but did not give up his former conception as regards 
the relationship of the three forms. In Coptotermes formosanus three types 
are also found, and each of these resembles rather closely the corresponding 
one in the Brazilian termite. The results of my studies of these forms have 
convinced me that Hartmann’s view is by no means right, and not only the 
“male” and the “female,” but also the “young” forms, should be referred 
to independent genera. 
Three more species of Pseudotrichonympha have been described recently. 
In his latest work (1917), Grassi described two new species of this genus, 
P. magnipapillosa and P. parvipapillosa, harboured by African (French 
Guinea) and Australian species of Schedorhinotermes. One more species was 
briefly described by Imms (1919) 2 from the Indian Archotermopsis . “ Tricho¬ 
nympha ” leidyi, described by Kent (1885), probably belongs to this genus; 
but his account is not sufficiently detailed to make its precise determination 
possible. In spite of the resemblances, the species of Pseudotrichonympha in 
our termites shows some differences from the Brazilian, Australian, African, 
and Indian species, so that it should be regarded as new, and I have therefore 
proposed the name Pseudotrichonympha grassii for it. A discussion of Hart¬ 
mann’s view of the relations between the three types will be given later. 
Pseudotrichonympha grassii sp. nov. (Plate XI, figs. 14-23). 
This is one of the largest of the protozoa found in our termites, reaching 
a size visible by the naked eye. The body usually assumes a spindle shape, 
measuring 200-300 p in length and 50-120 p in breadth. Sometimes it assumes 
a slender lanceolate shape, and its length exceeds 500 p in extreme cases. 
It is thus shorter and broader than Hartmann’s species and those described 
by Grassi. It is markedly variable in shape as compared with Trichonympha 
and other forms found associated in Coptotermes. 
In general organization it resembles Trichonympha, and the body may be 
distinguished into an anterior region or head, and a posterior region or body. 
1 Incorrectly called Captotermes by Hartmann. 
2 Incorrectly called “Trichonympha ( Holomastigotoides )” by him. 
