298 
Intestinal Protozoa in Termites 
respectively homologous with the wall of the axial core and its contents in 
Trichonympha. My knowledge of the process of division in this species is 
incomplete, but it seems highly probable that the slender strand and the 
structures acting as division-centres at both its ends are derived from the 
tubule. Each row of basal granules starts at the tip of the tubule, and the 
relation between them seems also analogous with what is seen in Trichonym'pha. 
There is, moreover, a structure which I conjecture to be homologous with a 
nuclear sac or corbule. This is the special protoplasm enveloping the tubule 
and the nucleus. The outer boundary of this layer of protoplasm is not 
distinctly contoured, but it is especially distinct at the base of the tubule 
and the anterior pole of the nucleus. As described in a previous section, the 
posterior end of the tubule is commonly found fixed on the nuclear wall; 
but sometimes they are separated from each other and connected by means 
of a mass of dense protoplasm, in which the contents of the tubule and the 
special protoplasm around it are not distinguishable. Thus I think there are 
many reasons indicating that the dense protoplasm and the contents of the 
tubule are composed of the same substance; and, moreover, that they are 
homologous with the nuclear sac, the corbule, and the contents of the axial 
core. From the above data, it may be concluded that the structure of Micro¬ 
spironympha is similar to that of Trichonympha, but of a less complex type. 
In Holomastigotoides , the organization of the anterior portion is apparently 
of a quite different type. There is no structure corresponding to the tubule 
in Microspironympha or the axial core in Trichonympha. All parts of the 
mitotic figure seem to be formed from the nucleus itself, and both the division- 
centre and the strand appear on the nucleus: so that it is reasonable to suppose 
that a body which acts as a division-centre is localized on the nuclear wall. 
The most interesting structure of Holomastigotoides, from the viewpoint of 
comparative morphology, is the peculiar mass of special protoplasm sur¬ 
rounding the nucleus and occupying the area in front of it. It is clearly ob¬ 
servable in good preparations that the anterior tip of this mass reaches the 
anterior extremity of the body, where the ends of the rows of basal granules 
of the flagella are kept in connexion with it. That is to say, in this genus 
the body playing the role of the division-centre lies separated from the basal 
granules of the flagella, but they are kept in connexion by means of the above- 
mentioned protoplasm. I believe that it is reasonable to interpret this 
structure as homologous with the dense protoplasm and the contents of the 
tubule of Microspironympha, and consequently with the corbule of Tricho¬ 
nympha and the nuclear sac of Teratonympha. 
In some individuals of Holomastigotoides the particular protoplasm is 
remarkably developed, assuming the form of a distinct and fairly long 
column hanging along the longitudinal axis of the body. In Microspiro¬ 
nympha some individuals are found provided with a structure at the posterior 
portion of the body, appearing sometimes as a well-developed transparent 
style. I am of the opinion that both these structures are similar in nature, 
