June, 2CC0 
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quivering Die female’s response is to 
become alert and approach. 
Wail 
This is a plaintive sounding soft cry 
made by the adult female during 
oestrous She makes it when 
approaching the courting male and 
when at the latrine during the courting 
period and after copulation It is made 
with the mouth slightly open and head 
thrust forward 
Gurring 
An often repeated single sound (every 
15 sec. to two nun.), it was made only 
by two adult females. It was made in 
bouts lasting over 15 minutes and 
always while resting. It may be the 
same sound as “bleating” observed by 
Milner & Harris ( 1999 ) It has no 
apparent stimulus 
Anticipatory or sucking grunt 
A low gruff sound, it is uttered by the 
male or female when approaching each 
other during courtship, investigating 
a new object, and by the mother while 
young are suckling. It is also made 
when anticipating offered food When 
uttered by mother suckling young it 
starts 10 sec. 3ftcr the beginning of 
suckling. The flanks heave with each 
sound and the body rocks forward and 
backward. 
Agonistic, alarm, tension or protest 
sounds 
Scream-yelp 
This sound is a raucous, harsh cry 
uttered by females protesting 
attempted mounting by the male, or 
when held in the hand It is apparently 
a startle response. 
Quack 
A sound like that made by a duck, this 
is made when one animal is being 
approached by another. It may be 
protest, startle or a defensive threat 
Creak 
A sound like an unoiled hinge, it is 
made in moments of uncertainty, as a 
protest or a slight threat. 
Hanh m iner 
This is like the soft twitter, but harsher. 
It is produced by juveniles when they 
move towards another animal or are 
approached by another. It causes the 
mother, or other animals, to become 
alert. It’s sonographic similarity to the 
screech indicates that it later develops 
into this call. 
Bark 
A single sound, it is repeated no more 
than tw ice and is in response to seeing 
a dog or a cat. It is a startle response 
and the animals become alert. 
Screech 
This is a harsh, loud, high-pitched 
sound, repealed several times, usually 
by a juvenile male during an aggressive 
interaction, in response to another 
animal’s screech, or by a female 
inspecting the primary home after 
anothet animal has visited it. Usually 
the animal screeching shows a high 
degree of excitement and rushes 
around. The response is alertness; 
juveniles approach the one making the 
