1(H) PROCEEDINGS: BOSTON SOCIETY NATURAL HISTORY. 
4. Next specimen. The posterior lobes are now abdominal, lying 
to the left of the intestine, which is displaced to the right and 
dorsally. The canals from the green glands reach the region of the 
pericardium but are not yet united. The sexual cells are abdominal. 
A rudimentary intestinal caecum is present. 
5. With livers completely shifted (pi. 8, fig. 40). The dorsal 
lobes have disappeared, and the anterior are greatly reduced. The 
chitinous gut is rapidly elongating, advancing the caecum towards 
the anterior part of the abdomen. The green glands have formed 
the nephrosac. The muscles are beginning to degenerate. 
6. Fourth day in the shell. The anterior lobes of the liver are 
gone. The ceils of the lateral caeca have taken on the adult his¬ 
tology and reactions; the nephrosac is abdominal; the muscles and 
pleopods and all the other organs are of adult type, in readiness for 
the moult to the adolescent phase. 
Sixth-stage crabs retain the simple cylindrical livers (pi. 8, fig. 42) 
for only a few days; then diverticula begin to appear along the 
borders, first of the right, then of the left gland (pi. 8, fig. 43). 
The development of these diverticula seems to follow a fairly definite 
plan (pi. 8, fig. 44). Owing to the way in which those from the 
right gland pass under the intestine, the adult condition is ultimately 
produced and the earlier displacement is obscured to casual inspec¬ 
tion. The livers seem to lie each on its own side of the intestine. 
A shift of the latter back again toward the mid-line of the body, 
which becomes possible from the eighth stage on because of the 
gradual proportional increase of the diameter of the abdomen, also 
aids in confusing the earlier relations. But by careful dissection, 
the displacement can be traced in the adult. The main axis of the 
right gland will be found to lie beneath or slightly to the left of the 
gut for a considerable part of its course in the abdomen. As young 
crabs show this better than older ones, two views of the abdominal 
contents of the crabs collected at Wareham are appended (pi. 8, 
figs. 45, 46). 
The green glands of JEupagurus longicarpus have the same gen¬ 
eral arrangement as the glands of A 7 , bernhardus described by 
Marchal (’ 92 ), except that the nephrosac is short and broad and the 
canals which unite this structure to the cephalic portion of the 
gland are without accessory diverticula. The cells of the canals 
and nephrosac have a characteristic histological appearance (pi. 9, 
