KNUTSON & GOSLINER: NEW SPECIES OF GTMNODORZS NUDIBRANCHS 
FROM THE PHILIPPINES 
135 
The overall body profile of G brunnea is 
similar to the speeies depleted as Gymnodoris 
marginata (Odhner, 1917). Gymnodoris mar- 
ginata also has gills arranged in a linear 
“bow.” A major differenee, however, is the 
position of the genital aperture, whieh is loeat- 
ed well forward of that in G brunnea. The 
radular teeth depleted in the G. marginata 
deseription are also very different in shape to 
those of G. brunnea, as the inner lateral of 
G marginata looks nearly identieal to the outer 
lateral depleted, whereas in G brunnea the 
first and seeond inner lateral teeth are elearly 
differentiated from the outer laterals. 
This speeies has only been doeumented 
from the Anilao region of the Philippines and 
it is unelear if it is restrieted to this region or 
has a wider distribution. We have a photo of a 
Gymnodoris speeimen from Japan that has a 
similar appearanee to G. brunnea. This speei¬ 
men appears to have a broader foot. It also 
does not seem to have the same dark brownish 
pigment to the skin and more subtle orange on 
the front margin of the foot. These animals 
would have to be further investigated via DNA 
or disseetions to eonfirm if they belong to the 
same or a different, elosely-related speeies. 
Gymnodoris tuberculosa Knutson and 
Gosliner, sp. nov. 
Figures 1 C-D, 5, 6, 7 
Gymnodoris sp. 3. Gosliner, et al, 2008: 149. 
Type Material. — Holotype: CASIZ 
191190, one speeimen, “Kranket Fish Market 
Cove,” near Madang Resort, Madang Prov- 
inee, Papua New Guinea, 5°12'28.4148"S, 
145°4831.9896"E, eolleeted at night, Novem¬ 
ber 13, 2012 by Vanessa Knutson, preserved 
speeimen 6.0 mm length. Paratypes: CASIZ 
083765, one speeimen, disseeted, 18.6m 
depth, Bonito Island, Batangas, Luzon Island, 
Philippines, eolleeted February 22, 1992 by T.M. Gosliner, preserved speeimen 8.3 mm length. 
CASIZ 086418, one speeimen, disseeted, 5.2 m depth, S. side of Raseh Passage, Madang, Papua 
New Guinea, eolleeted June 17, 1992 by T.M. Gosliner, preserved speeimen 11.0 mm length. 
CASIZ 177268, one speeimen, 14.2 m depth, Layag Layag, Caban Island, Marieaban Island, 
Batangas Provinee, Luzon, Philippines, eolleeted Mareh 16, 2008, preserved speeimen 4.0 mm 
Figure 4. Light micrographs of the penial spines: seale 
bar=100 pm. A. G. brunnea sp. nov., CASIZ 185973B. G. 
tuberculosa sp. nov., CASIZ 185954. C. G pseudobrunnea 
sp. nov., CASIZ 185967. 
