KNUTSON & GOSLINER: NEW SPECIES OF G TM7VOZ) ORZS NUD IB RANCH S 
FROM THE PHILIPPINES 
137 
embedded in the skin. The spicules can be seen through the skin in some of the photos and are pres¬ 
ent in the preserved specimens. These spicules vary greatly, in one specimen, CASIZ 191461 there 
are spicules that have a pink tinge to them while others appear colorless. There also appears to be 
different morphologies for the spicules within a specimen. Some of the larger spicules were “T” or 
“Y”-shaped and approximately 0.2 mm long, others were smaller and rod-shaped. 
Internal Anatomy.— On the ventmm and sides of the animal there are slightly branched, 
thickened glandular structures (Fig. 5). These structures are white to pale yellow in color and 
appear to originate at the foot and come up the sides of the animal. Although these stmctures are 
subcutaneous, they are clearly visible through the translucent skin in the living animal and in pre¬ 
served specimens. These stmctures seem to be concentrated anterior to and at the level of the gills. 
The stmctures do not appear to be present in the posterior portions of the foot of the living animals. 
These stmctures are more pronounced in some specimens than in others. They resemble the sub¬ 
cutaneous glandular tubes described for Gymnodoris aurita (Gould, 1852), (Gosliner 1997), and 
presumably serve a defensive function. 
Buccal and radular morphology: The buccal mass is muscular and has long strap-like salivary 
glands. Typical of the genus, the radula lacks a rachidian row of teeth (Fig. 6A). The inner lateral 
tooth (Fig. 6B) is a curved hook with a thickened, relatively straight base. The mid lateral teeth 
(Fig. 6C) are hook shaped with an elongate cusp. The outermost laterals (Fig. 6D) are thinner than 
the mid laterals and have a less pronounced base and a more elongate cusp. The radular formulas 
of four specimens were counted as follows: 20x20.1.0.1.20 (CASIZ 185954), 21 x21.1.0.1.21 
(CASIZ 191518), 21 X 20.1.0.1.20 (CASIZ 083765), 18 x 15.1.0.1.15 (CASIZ 086418). 
Reproductive system: There 
are two ovotestis masses located 
anterior to the digestive gland 
and posterior to the genital mass. 
A hermaphroditic duct extends 
from each of the ovotestis folli¬ 
cles (Fig. 7). Just anterior to their 
junction, the hermaphroditic duct 
widens into the ampulla. The 
ampulla then bifurcates into a 
short oviduct which enters the 
female gland mass. The second 
branch widens into the curved 
prostate, which surrounds the 
bursa copulatrix and then nar¬ 
rows into the muscular vas defer¬ 
ens. The vas deferens consists of 
several convolutions and coils 
before entering the penis, adja¬ 
cent to the vagina. The uterine 
duct emerges from the female 
glands near the oviduct and 
enters the small pyriform recep- 
taculum seminis. The elongate 
receptaculum duct enters the 
vagina at the beginning of a 
Figure 6. Gymnodoris tuberculosa sp. nov. scanning electron micrographs 
of the radula. A. View of the entire radula, CASIZ 185954, seale= 90 pm. 
B. Inner lateral teeth, CASIZ 185954, seale= 20 pm. C. Mid lateral teeth, 
CASIZ 185954, seale= 20pm. D. Outer laterals, CASIZ 086318 seale= 20pm. 
