140 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
bursa copulatrix and narrows into 
the muscular vas deferens. The 
vas deferens consists of a convo¬ 
lution where it widens and enters 
the penis, adjacent to the vagina. 
The uterine duct emerges from 
the female gland mass and joins 
with the pyriform receptaculum 
seminis. The receptaculum duct 
enters the vagina midway 
between the bursa copulatrix and 
the genital aperture. The albumen 
and membrane glands are small 
and not well differentiated. The 
mucous gland comprises the 
majority of the female gland 
mass. The penis of one specimen 
(CASIZ 185967) was cleared and 
stained (Fig. 4C) and is armed 
with densely distributed spines 
that have a broad base (roughly 
19pm across). The spines appear 
to be longer at the base of the 
penis, roughly 67 pm, and short¬ 
er at the very tip, roughly 31 pm. 
Natural history.— This 
species was collected at night 
from a sandy bottom and likely 
burrows in the sand. One of the specimens dissect¬ 
ed (CASIZ 185967) had a minimally digested nudi- 
branch in its stomach. The nudibranch pulled from 
the stomach had a dark coloration concentrated on 
the dorsal side, consistent with G. brunnea. After 
imaging the radula of this nudibranch (Fig. 11) with 
a scanning electron microscope, it is clear that it is 
consistent with that of G. brunnea with large, hook¬ 
shaped inner laterals and distinctive bicuspid sec¬ 
ond lateral teeth. Due to the early stage of digestion 
of the stomach contents, it is possible that feeding 
took place following collection, if the two speci¬ 
mens were placed in the same container at the time 
of collection. If this is what occurred, one may 
Figure 9. Gymnodoris pseudobrunnea sp. nov. scanning electron micro¬ 
graphs of the radula. A. View of the entire radula, CASIZ 185969, seale = 60 
pm. B. Inner and mid lateral teeth, CASIZ 185969, seale = 30 pm. C. Outer 
lateral teeth, CASIZ 185969, seale = 30 pm. 
Figure 10. Gymnodoris pseudobrunnea sp. nov. reprodue- 
tive system: seale bar = l mm. Abbreviations: am, ampulla; be, 
bursa eopulatrix; fgm, female gland mass; p, penis; pr, prostate; 
rs, reeeptaeulum seminis; v, vagina; vd, vas deferens. 
