142 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
The morphological differences show clearly that these are two different species, and the 
molecular data presented here further corroborate their distinctiveness. The uncorrected COI p-dis- 
tances within G brunnea specimens ranged from 0.0% to 0.5%, representing intraspecific diversi¬ 
ty. We were only able to successfully amplify the COI fragment for one specimen of G. pseudo- 
brunnea and, although with only one specimen we cannot measure any intraspecific genetic vari¬ 
ation for this species, this one specimen ranges from 10.3-11.1% different in the COI fragment 
from the G. brunnea specimens. This is well in line with the mean COI divergence of Herbert et al. 
(2003), which found a mean divergence of 11.1 ±5.1% for congeneric pairs within Mollusca, and 
above the 5.5% minimum p-distance cutoff applied in a recent paper that delineated sister species 
of aeolid nudibranchs (Carmona et al. 2013). 
Acknowledgments 
We would like to thank Michelle Weber for her excellent Gymnodoris spotting skills and Alex¬ 
is Principe, Peri Palaracio and Ditto de la Rosa for being fantastic dive guides and for their talent 
of finding interesting specimens. We also acknowledge Will and Margaret Hearst for the funds to 
be able to survey this region of the Philippines, as well as BEAR for providing us with the permits, 
and the mayor of Mabini for supporting our efforts. We would further like to acknowledge the ‘Our 
Planet Reviewed’ Initiative, Papua New Guinea 2012-2103 Expedition and Philippe Bouchet for 
including the first author as a part of the shallow water marine portion of the expedition. Further 
thanks to all the colleagues who have contributed specimens over the course of this project and to 
the reviewer for suggestions on how to improve the manuscript. 
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