216 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
Lissodiadema foriofi 
.‘(lUf’u'l 
Cieeirt 
Cfypeaster reticufatus 
jeotiii* 
□ eian 
Figure 2. Four exemplars of major Philippine eehinoid taxa and their eorresponding distributions. Seale bars in animal 
images are 10 mm long. Range maps are edited from the Eneyelopedia of Life (EoL), with red dots indieating EoL reeords 
as well as new data from the 2011 Hearst Philippine Biodiversity Expedition. 
throughout the Indo-Pacific. Our list includes 210 species so far verifiably recorded from the 
Philippine region (see Appendix). We regard this estimate of eehinoid diversity in the Philippines 
as conservative, given that there are several undescribed species of clypeasteroids and spatangoids 
either listed in van Noordenburg (2008) or presently being worked on by the senior author and his 
colleagues. We also do not separately enumerate subspecies, some of which might turn out to be 
full species. Finally, we note that there will likely be new material coming out of work such as that 
described by Bouchet et al. (2004), potentially adding to our compilation. 
To build the list, we relied on a variety of literature and collections-based sources. In his mon¬ 
umental Monograph of the Echinoidea, Mortensen (1928, 1935, 1940, 1943a, 1943b, 1948a, 
