258 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
spot; blackish spots or dashes on 
top of snout and interorbital 
spaee, ineluding dorsal aspeet of 
eornea of eye. Dark brownish 
spots on oeeipit immediately 
behind orbits. Mouth and gill 
eavities pale. First dorsal fm 
blaek exeept pale at immediate 
base of first spine and distal tip 
of same. All other fins without 
pigmentation. 
Etymology.— It is a pleas¬ 
ure to name this fish after Alan E. 
Leviton, Curator Emeritus of the 
California Aeademy of Seienees, 
who strongly promoted this 
Hearst Expedition volume, used 
his eonsiderable teehnieal knowl¬ 
edge of desk-top publishing to 
produee this work, and who pro¬ 
vided mueh adviee and support with this paper. 
Remarks and comparisons. — These speeimens are elosely similar in most eharaeters to 
P. natalensis, with only a few morphometrie differenees noted in eaudal-pedunele depth, and pee- 
toral and pelvie fm lengths (Table 3). The type speeimens of P. natalensis were not in the best 
shape: fins were damaged and not fully intaet, and no seales or eolor pattern remained. Three other 
speeimens reeorded by Smith and Johnson (2007:374) have provided information on lengths of 
median-fin rays and expanded the range of eounts of seeond dorsal-fin and anal-fin rays. The 
HEPD speeimens are, in all likelihood, subadults and thus may not display features eharaeterizing 
mature individuals. 
Figure 6. Pteropsaron levitoni sp. nov. Photographs of cleared-and-stained 
specimen, CAS 236402 (24.5 mm SL). (a) dorsal view; (b) lateral view); (c) 
ventral view. 
Discussion 
The genera of hemeroeoetine Triehonotidae have been inadequately defined and a further shuf¬ 
fling of taxa within this group is likely, but it is beyond the seope of this paper to attempt sueh an 
undertaking. Nelson (1982:7) reeognized a elose relationship among the three genera, Acan- 
thaphritis, Pteropsaron, and Osopsaron, and diseussed the relationships of that elade to other per- 
eophids. Smith and Johnson (2007) further developed his eoneept of relationships and removed the 
Hemeroeoetinae and Creediidae from the Pereophidae and reeognized them as subfamilies in the 
expanded family Triehonotidae. 
That three different species of Hemeroeoetinae, two of whieh are new, were eolleeted in a sin¬ 
gle haul of a beam trawl in the Philippines makes it appear likely that many more speeies will be 
diseovered in the Indo-West Paeifie with more eolleeting at appropriate depths and bottom substra¬ 
tum. Beeause of their small adult size, eylindrieal body shape, and probable eryptie sand-dwelling 
habits, they have avoided eapture and are possibly mueh more abundant and wide-spread than is 
apparent from museum speeimens alone. Current developing efforts in eolleeting in deeper waters 
with SCUBA should add markedly to the number of known speeies in this group. 
