IWAMOTO & MCCOSKER: DEEP-WATER FISHES OF THE PHILIPPINE EXPEDITION281 
and C. tydemani, but can be distinguished from them by a combination of the shape of its preopercle, 
length of elongated ID ray, orbit-to-preopercle distance, and the relationship of that measurement to size 
of its orbit (see key in Shcherbachev and Iwamoto 1995:286-287). 
Hymenocephalus lethonemus Jordan and Gilbert, 1904 
Material.— CAS 233709 (2 spec., 23.8-24.0 mm HE, 112-122 mm TL); HEPD-006, otter trawl in 454-157 
m. 
Remarks. — Distributed from southern Japan, Kyushu-Palau Ridge, East China Sea, South China Sea off Tai¬ 
wan to the Philippines. 
Hymenocephalus nascens Gilbert and Hubbs, 1920 (Fig. 56) 
Material.— CAS 233942 (85+ mm TL, DC-1288); HEPD-005, beam trawl in 459^96 m. 
Remarks. — This species is closely similar to H. lethonemus. It has been recorded from the Philippines to the 
southwestern Pacific off New Caledonia and Australia (QLD, NSW, WA), in about 350-930 m. 
Hymenocephalus torvus Smith and Radcliffe in Radcliffe, 1912 
Material.— CAS 234080 (7, 49+-113i-om TL); HEPD-009, otter frawl in 322-336 m. CAS 234076 (9, 
21-27 mm HL, 75+-146 mm TL); HEPD-010, beam trawl in 318-333 m. CAS 234083 (2, 24.5-27.8 mm 
HL, 370+-370+ mm TL); HEPD 035, beam trawl in 397-439 m. CAS uncat (73.5 mm HL, 287+ mm TL); 
HEPD-012, beam Lawl in 721-761 m. 
Remarks.— -This species was recognized as a subspecies of H. striatissimus by Gilbert and Hubbs 
(1920:530), but the V-ray count, barbel length, and body pigmentation appear to distinguish the two 
species. According to Gilbert and Hubbs (1920: 528) the nominal subspecies ranges from Japan “south¬ 
ward to China, Formosa, and the east coast of Luzon,” “intergrading off the northwest coast of Luzon with 
the subspecies tonms, which inhabits the Sulu Sea and the China Sea off southern Luzon.” Those authora 
coiTected the Albatross station number (sta. 5508) for the holotype ofi/. torvus, erroneously given by Rad¬ 
cliffe (1912:1 10) as station 5548. 
Mataeocephalus acipenserinus (Gilbert and Cramer, 1897) 
Material.— CAS 235251 (51.2 mm HL, damaged); HEPD-011, otter trawl in 636-664 m. CAS 235822 (47 
mm SL, 225+ mm TL); HEPD-39, beam trawl in 633-642 m. 
Remarks. — Widely distributed in tire tropical Indo-Pacific, from the Sala-y-Gomez Ridge in the southeast¬ 
ern Pacific to Hawaii, southwestern Pacific, Phihppines, Indonesia, and South China Sea, across Indian 
Ocean to east Africa. 
Mataeocephalus cristatus Sazonov, Shcherbachev and Iwamoto, 2003) (Fig. 57) 
Material.— CAS 234020 (46.2 mm HL, 210 mm TL, DC-1383); HEPD-012, beam trawl in 721-761 m. 
Remarks. — This species is known from the Indian Ocean (Saya de Malha Ridge, Mascarene Ridge, Ninety- 
East Ridge), the South China Sea (Paracel Is.), and the Markus-Necker Ridge in the western Pacific. The 
HEPD specimen is the first record from the Pliilippines. 
Nezumia spinosa (Gilbert and Hubbs, 1916) (Fig. 58) 
Material.— CAS 235798 (44.6 mm HL, 220+ mm TL, DC-1442); HEPD-019, beam trawl in 541-636 m. 
Remarks.— A widespread species known from Japan south and east to Australia (QLD and WA) and New 
Caledonia, and west to southeastern Africa. 
Sphagemacrurus pumiliceps (Alcock, 1894) (Fig. 59) 
Material.— CAS 235732 (29.0 mm HL, 1 87+ mm TL, DC-1412) and CAS 236022 (28.2 mm HL, 190 mm 
TL); HEPD-01 6 , otter trawl in 892-966 m. 
Remarks. — Apparently widespread tliroughout Indo-West Pacific, but more than one species may be liiding 
under this name. 
