IWAMOTO & MCCOSKER: DEEP-WATER FISHES OF THE PHILIPPINE EXPEDITION297 
Bothidae (Lefteye Flounders) 
Arnoglossus elongatus Weber, 1913? 
Material.— CAS 235544 (2, 85-95 mm SL); HEPD-021, beam trawl in 132-172 m, 
Remarks.— These two HEPD specimens key out rather readily in Hensley and Amaoka’s (2001) key to the 
species of Arnoglossus. Closer comparisons with other specimens, especially males, must be made before 
a more-definitive identification can be made. The species is known from Taiwan, Philippines, Madura (in 
Java Sea), and northwestern Australia, in 100-224 m. 
Arnoglossus japonicus Hubbs, 1915 (Fig. 145) 
Material.— CAS 235577 (13, 53-101 mm SL) and CAS 2355786 (127mm SL, DC-1499); HEPD-022, 
otter trawl in 115-144 m. 
Remarks.— Known distribution from southern Japan to Gulf of Tonkin and New Caledonia; the HEPD spec¬ 
imens represent the first record from the Philippines. Attains 17 cm TL. 
Arnoglossus oxyrhynchus Amaoka, 1969 (Fig. 146) 
Material.— CAS 235549 (150 mm SL, DC-1528); HEPD-030, beam trawl in 190-201 m. 
Remarks.— Possibly a junior synonym of .4. bninneus (Fowler, 1934). It occurs in depths of about 100-300 
m and is known from southern Japan (Kochi), with an uncon fiiTncd record from New Caledonia. Arnoglos¬ 
sus bninneus is known only from the east coast of Luzon in 292 m. (Hensley and Amaoka, in Carpenter 
and Niem, 2001:3827) 
Kamoharaia megastoma (Kamohara, 1936) (Fig. 147) 
Material.— CAS 235542 (89 mm SL, DC-1473); HEPD-021, beam trawl in 132-172 m. CAS 235535 (88.5 
mm SL); HEPD-022, otter trawl in 115-144 m. 
Remarks. — Previously known from southern Japan to Taiwan and in the In do-Australian Archipelago in 
300-500 m. Tlic HEPD specimens are the first recorded from the Phihppines, so far as we know. 
Lae ops clarus Fowler, 1934 
Material.— CAS 235744 (125.2 mm SL, DC-1468); HEPD-021, beam trawl in 132-172 m. 
Remarks.— This specimen appears to be very similar to Laeops sp. from HEPD-036, agreeing rather well in 
mcristic values and most other characters. However, the body is deeper, the eyes larger and have broad 
black margins along the mesial and posterior borders (light to medium brown in HEPD-036), and the bran- 
chiostegal membrane is black (compared with medium brown).The HEPD specimen may be only the sec¬ 
ond record of the species. 
Laeops sp. 
Material.— CAS 236558 (2, 106-120 mm SL); HEPD-036, beam trawl in 216-262 m. 
Remarks. — These specimens agree most closely with the Japanese species L. kitaharae (Smitli and Pope, 
1906) except for the deeper body (<3 time in SL) and the presence of fine spots in that species. Laeops 
clarus Fowler, 1934, apparently known only from the Phihppines, has a slender body but the HL goes 5.7 
times in SL and there are 10 GR on the lower limb of the first arch. Laeops gixicilis Fowler, 1934, also 
known only from the Philippines, has higher counts of rays in its D (112) and A (93) and the HL goes about 
five times in SL. 
Parabothus taiwanensis Amaoka and Shen, 1993 (Fig. 148) 
Material.— CAS 235505 (107 mm SL, DC-1510); HEPD-026, beam trawl in 82-86 m. 
Remarks.— Tliis specimen may represent the first record of the species outside Taiwan. Anaoka and Shen 
(1993) describe the teeth as biserial in its upper jaw, but that does not appear to be the case in the HEPD 
specimen; also they describe the body as pale puiplish after removal of scales (vs. cream-colored). Other¬ 
wise the description fits our specimen rather well. 
