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THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
Figure 12. Internal anatomy oiPhiline multipapillata sp. nov., holotype, CASIZ 172841. A. Scanning eletron micro¬ 
graph of unpaired gizzard plate. B. Scanning electron micrograph of paired gizzard plate. C. Scanning electron micrograph 
of gizzard plate micro structure. D. Scanning electron micrograph of half-row of radular teeth. E. Dorsal light micrograph 
of penis. F. Ventral light micrograph of penis. 
Digestive system (Fig. 12A-D, 13B): The buccal mass is relatively small, occupying the ante¬ 
rior quarter of the anterior shield. The buccal bulb has a relatively small radula. The radular formu¬ 
la in the holotype is 20 x 1.0.1. The inner lateral teeth (Fig. 12D) are large and broad with a curved 
prominent cusp. Their masticatory margin has a series of up to 56 elongate, closely crowded den¬ 
ticles. Outer lateral teeth are absent. There is a large ventral oral gland and small dorsal oral glands. 
At the posterior end of the buccal mass, near the junction with the crop, is a pair of short salivary 
glands. The gizzard is large and highly muscular, much wider than the buccal bulb. There are 
unequal gizzard plates (Fig. 13B) that are elongate with rounded apices. The unpaired plate is 
smaller (Fig. 12A) than the paired plates (Fig. 12B). All plates have paired holes that are elongate 
and large in size and are relatively equal in size. There is an organized pattern of honeycombed fine 
