New Ray Spiders from Southeast Asia: 
The New Philippine Genus Tagalogonia gen. nov. and 
Continental Genus Coddingtonia Miller, Griswold and Yin, 2009 
(Araneae: Theridiosomatidae), with Comments on Their 
Intergeneric Relationships 
Facundo M. Labarque and Charles E. Griswold 
^ Arachnology Lab, Entomology' Department, California Academy of Sciences, 55 Music Concourse 
Drive, San Francisco, CA 94118, USA; ~ Biolog Department, San Francisco State University, 
1600 Holloway Ave, San Francisco, CA 94132, USA: ^ Environmental Science, Policy and 
Management, University of California, 101 Sproul Hall, Berkeley, CA 94704, USA; ^ Biolog 
Department, The George Washington University, 21211 St NW, Washington, DC 20052, USA. 
* Address correspondence to facundo.labarque@gmail.com 
A new genus of Theridiosomatidae from the Philippines is described based on male 
and female characters: Tagalogonia gen. nov. Two new species are described for the 
genus: Tagalogonia banahaw sp. nov. and Tagalogonia isarog sp. nov. Anew species of 
Coddingtonia Miller, Griswold and Yin, 2009 is described based on males and females 
from Malaysia: C anaktakun sp. nov. The monotypic genus Luangnarn Wunderlich, 
2011 is considered a junior synonym of the monotypic Coddingtonia syn. nov. The 
female of Coddingtonia discobulbus (Wunderlich, 2011) comb. nov. is illustrated. Dis¬ 
cussions on the intergeneric relationships of the Asian Theridiosomatidae are given. 
Keywords: Symphytognathoids, Indomalaya Region, taxonomy, new species, new syn¬ 
onymy, Luangnarn. 
The ray spiders, family Theridiosomatidae, are membei*s of a monophyletic group of mostly 
minute orb-weavers (Griswold et al. 1998; Schiitt 2003; Lopardo et al. 2010). Theridiosomatidae 
area diverse group, currently with 18 genera and 101 species distributed mostly in tropical areas all 
around the world (Platnick 2014). The family has been reviewed worldwide at the generic level 
(Coddington 1986), with recent contributions of additional new species from South America 
(Rodrigues and Ott 2005a-b; Rodrigues and Lise 2008) and Asia (Miller, Griswold and Yin 2009; 
Ono and Shinkai 2009; Shinkai 2009; Chen 2010; Wunderlich 2011; Dou and Lin 2012; Zhao and 
Liu 2012). 
Several new genera were described from Asia including Karstia Chen, 2010, Chthonopes 
Wunderlich 2011, the monotypic Coddingtonia Miller, Griswold and Yin 2009, Luangnarn Wun¬ 
derlich 2011, Alaria Zhao and Liu 2012 and Menglunia Zhao and Liu 2012. The main diagnostic 
character that defines Coddingtonia, the separated spennathecae (Miller et al. 2009), is also shared 
by Chthonopes, Luangrram and Menglunia (figs. 7, 18f Wunderlich 2011; figs. 16B, 18D Zhao and 
Liu 2012). In the presence of new evidence, the genus Luangnarn is here considered a junior syn¬ 
onym of Coddingtonia (see Systematics and Taxonomy). Coddingtonia was fnst described based 
on only one female specimen, and Luangnarn was described based in several male and female spec¬ 
imens. However, the Luangnarn female illustrations are not detailed enough to provide conclusive 
evidence of validity. The discovery of a new species of Coddingtonia from Malaysia represented 
by specimens of both sexes (see Taxonomy) was the key to synonymize Luangnarn. Coddingtonia 
discobulbus (Wunderlich, 2011) comb. nov. is redescribed and the female illustrated based on sev- 
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