LABARQUE & GRISWOLD: NEW RAY SPIDERS FROM SOUTHEAST ASIA 
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five years, many new genera and species of Theridiosomatidae were described for Asia (Miller, 
Griswold and Yin 2009; Ono and Shinkai 2009; Shinkai 2009; Chen 2010; Wunderlich 2011; Dou 
and Lin 2012; Zhao and Liu 2012), but the intergeneric and subfamilial relations of those genera 
were not integrated into the phylogenetic framework proposed by Coddington (1986). Here we 
present a modified version of the diagnostic characters of Theridiosomatidae genera (fig. 1 Cod¬ 
dington 1986) including all the new genera and several new diagnostic characters (Table 1). 
Andasta Simon, 1895, Karstia Chen, 2010, and Zoma Saaristo, 1996 share the characters that 
defined Ogulninae and Theridiosomatinae; the cymbial lamella, the embolic apophysis, the embo¬ 
lus as a thin lamina, the basal sperm duct opening, the embolic division covered by the conductor, 
and the convoluted copulatoiy ducts (figs. 152-156 Coddington 1986; figs. 6, 11-13, 20, 25-27 
Chen 2010; figs. 28A-D, 29B Zhao and Liu 2012) and, especially the male charactei*s synapomor- 
phic for Theridiosomatinae including the striated tegulum, the setae row on the cymbial lamella, 
and the elongated median apophysis with a median gi*oove (figs. 133, 153, 163, 186, 197, 211 Cod¬ 
dington 1986; figs. 10, 24 Chen 2010; figs. 28A-D Zhao and Liu 2012). Chthonopes Wunderlich, 
2011, also share most of those characters, except the setae row on the cymbial lamella (figs. 13-14 
Wunderlich 2011). Because of those synapomoiphies we here include Andasta Simon, Chthonopes 
Wunderlich, Karstia Chen, and Zoma Saaristo in the subfamily Theridiosomatinae (Table 1). 
Andasta and Theridiosoma, and Karstia and Baalzebiib share the same diagnostic characters 
respectively, and thus are considered closely related to each other (Table 1). Zoma seem to be close¬ 
ly related to Andasta and Theridiosoma having a striated tegulum, a tegular spur, a setae row on 
the cymbial lamella, an epigynum with lateral pits, a median apophysis with a dorsal median 
groove and copulatory ducts inserted mesally on the spermathecae (figs. 28A-D, 29A-B, 30A-D 
Zhao and Liu 2012; figs. 145-149, 151-156 Coddington 1986). Zoma also share with Epilineutes 
and Wendilgarda the stripes on the conductor (figs. 28C-D, 30B Zhao and Liu 2012; figs. 189, 197 
Coddington 1986). Chthonopes seem to be closely related to Epilineutes and Wendilgarda having 
a sub-rectangular median apophysis with denticles and an epigynal scape (figs. 17,18b, 18e Wun¬ 
derlich 2011; figs. 173, 183, 189, 197 Coddington 1986). Unfortunately, several diagnostic charac¬ 
ters are not well documented in Chthonopes (Table l;Wunderlich 2011). On the other hand, Zoma 
and Chthonopes share with Platoninae, Epeirotypinae, Menglunia, Ogidninae, Coddingtonia and 
Tagalogonia gen. nov. a transverse posterior groove on the epigynum (Figs. IB, D, 3D-F, 6D-F, 
7D-F; figs. 25, 37, 50, 78 Coddington 1986; fig. 18f Wunderlich 2011; figs. 16B, 18D, 29B Zhao 
and Liu 2012), whereas this is absent in the other Theridiosomatidae. 
Coddingtonia and Tagalogonia gen. nov. share with Theridiosomatinae and Ogulninae the 
cymbial lamella, the embolic apophysis, the embolus as a thin lamina, the basal sperm duct open¬ 
ing, the embolic division covered by the conductor, and the convoluted copulatory ducts (Figs. 
lA-D, 2D-F, 3D-F, 4D-F, 5D-F, 6D-F, 7D-F; figs. 13-14, 18, 18c, 18f Wunderhch, 2011; figs. 
102, 113, 152-153, 164, 184, 191, 211, 215 Coddington 1986; figs. 6, 11, 20, 25 Chen, 2010; figs. 
28A-D, 30C-D Zhao and Liu, 2012), and with Theridiosomatinae the mesal insertion of the cop¬ 
ulatory ducts on the spermathecae (Figs. IB, D, 3D, 6E, 7E; figs. 145,152,174,184,214-215 Cod¬ 
dington 1986; figs. 6, 20 Chen 2010; figs. 29B, 30D Zhao and Liu 2012), whereas other Theridio¬ 
somatidae lack the embolic division and its apophysis, the tube like embolus has a distal opening, 
the copulatory ducts are short and make a simple, acute turn, and the copulatory ducts insert later¬ 
ally. Coddingtonia and Tagalogonia gen. nov. seem to be closely related to Theridiosomatinae 
(Table 1), especially Coddingtonia that have a striated tegulum (Fig. 1C). However, both genera 
lack Theridiosomatinae synapomoiphic male characters such as the setae row on the cymbial 
lamella and the median apophysis with a dorsal median groove (Figs. 2D-F, 4D-F, 5D-F; figs. 3-5 
Wunderlich 2011; figs. 133, 153, 163, 186, 197,211 Coddington 1986). 
