412 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
Coddingtonia and Tagalogonia gen. nov. can be distinguished from other genera of Theridio- 
somatidae by the opisthosoma sparsely covered with long and strong setae, the well-developed disk 
shaped conductor, the long and slender cymbium with a ventral groove where the conductor fits, 
and the long and thin copulatory ducts (Figs. lA-D, 2A-F, 3A-F, 4A-F, 5A-F, 6A-F, 7A-F; figs. 
8B, llE-F Miller et al. 2009; fig. 3-7 Wunderlich 2011). Particularly, males of Coddingtonia and 
Tagalogonia gen. nov. resemble those of Baalzebub, Epilineutes, Karstia and Wendilgarda (all of 
them in Theridiosomatinae) by having a tripartite embolic apophysis (Figs. 1 A, C, 2D-F, 4D-F, 
5D-F; figs. 161-162, 191, 198 Coddington 1986; figs. 11, 25 Chen 2010;), whereas OgulniushdiYQ 
the embolic apophysis bifurcated (fig. 100 Coddington 1986) and Theridiosoma, Zoma andAndas- 
ta have it fragmented (fig. 133 Coddington 1986; fig. 28D Zhao and Li, 2012). The embolic apoph¬ 
ysis is not well documented in Chthonopes ffigs. 18, 18c Wunderlich, 2011). However, Codding¬ 
tonia and Tagalogonia gen. nov. lack the characters that support the clade of Epilineutes and 
Wendilgarda, e.g., the epigynal scape and the sub-rectangular median apophysis with denticles 
(figs. 173, 189, 197,213 Coddington 1986). On the other hand, females of Coddingtonia and Taga¬ 
logonia gen. nov. share with those of Platoninae, Epeirotypinae, Ogulninae, Chthonopes, Menglu- 
nia and Zoma the presence of a transverse posterior groove on the epigynum (Figs. IB, D, 3D-F, 
6D-F, 7D-F; figs. 25, 37, 50, 78 Coddington 1986; fig. 18f Wunderlich 2011; figs. 16B, 18D, 29B 
Zhao and Liu 2012), whereas this character is absent in the other Theridiosomatidae. Peculiarly, 
females of Coddingtonia and Tagalogonia gen. nov. share with those of Chthonopes and Menglu- 
nia the separated spermathecae (Figs. IB, C, 3D-F, 6D-F, 7D-F; fig. 18f Wunderhch 2011; figs. 
16B, 18D Zhao and Liu 2012), whereas these are juxtaposed or partially fused in the other 
Theridiosomatidae (Coddington 1986; Miller et al. 2009; Zhao and Liu 2012). 
Alaria and Menglunia resemble Platoninae and Epeirotypinae having the embolus tube-like 
with a distal opening, and lacking the embolic division and apophysis (figs. lA-D, 5A-B, 15A-B, 
17A-B, 18A-B Zhao and Liu 2012). Menglunia also share with Platoninae and Epeirotypinae short 
copulatory ducts with a simple and acute turn (figs, 16B, 18D Zhao and Liu 2012), whereas Alar¬ 
ia share with Coddingtonia, Tagalogonia gen. nov., Theridiosomatinae and Ogulninae convoluted 
copulatory ducts (figs. 2A, 5D Zhao and Liu 2012). Females of Alaria (figs. 2A-B Zhao and Liu 
2012) also share an epigynal scape with Chthonopes, Epilineutes and Wendilgarda (figs. 18d-e 
Wunderlich 2011; figs. 173, 213 Coddington 1986), while Alaria males lack all the Theridioso¬ 
matinae synapomorphies. Zhao and Liu (2012) suggested that Alaria is closely related to 
Chthonopes and Wendilgarda, but this seems unlikely. Taking into account all the former evi¬ 
dences, Menglunia and Alaria seems to be closely related to Platoninae and Epeirotypinae 
(Table 1). 
Taxonomy 
Genus Tagalogonia Labarque and Griswold, gen. nov. 
Type Species. — Tagalogonia banahaw Labarque and Griswold, sp. nov. 
Etymology.— The generic name is a noun in apposition that resembles the Austronesian lan¬ 
guage spoken as a first or second language in Philippines: Tagalog. The gender is feminine. 
Monophyly. — Putative synapomorphy: Males of Tagalogonia gen. nov. have a median 
apophysis very small (minute), square and smooth (Figs. 1 A, 2F, 4F). 
Diagnosis. — Tagalogonia gen. nov. and Coddingtonia can be distinguished from other gen¬ 
era of Theridiosomatidae by the opisthosoma sparsely covered with long and strong setae, the well- 
developed disk shaped transparent conductor, the long and slender cymbium with a ventral groove 
where the conductor fits, and the long and thin copulatoiy ducts (Figs. lA-D, 2A-F, 3A-F, 4A-F, 
5A-F, 6A-F, 7A-F; figs. 8B, 1 lE-F Miller et al. 2009; figs. 3-5 Wunderlich 2011). Males of Taga- 
