LABARQUE & GRISWOLD: NEW RAY SPIDERS FROM SOUTHEAST ASIA 
419 
DiSTRrouTiON. — Philippines, Luzon Island, Camarines Sur Province, Mt. Isarog; known only 
from the type locality. 
Genus Coddingtonia MUler, Griswold and Yin, 2009 
Coddingtonia Miller, Griswold and Yin, 2009: 30, figs. 8B, llE-F, 94. Type species Coddingtonia eury- 
opoides Miller, Griswold and Yin [one $ holotype (CASENT 9022403, HNU), one immature paratype 
(CASENT 9029336), from China, Yunnan Province, Longling Co., Mangkuan Township, Zaotang He at 
Baihualing village, examined] 
Luangnam Wunderlich, 2011: 431. Type species by monotypy Luangtiam discobulbus Wunderlich [one S 
holotype (SMF), one $ paratype (SMF), one and 5 $ paratypes (SMF-62412), from Laos, Luang Nam 
Tha Province, Vieng Phou Kka, Phou Pasat, Tliam Pasat Thia, examined] syn. nov. 
Synonymy.— The female of Luangnam discobulbus Wunderlich agrees in every way with the 
diagnostic characters here described for Coddingtonia Miller, Griswold and Yin, 2009, and the 
male agrees in detail with the diagnostic characters of males of our new species, Coddingtonia 
anaktakun Labarque and Griswold. 
Monophyly. — Putative synapomorphies: Males of Coddingtonia have a median apophysis 
elongated, distally acute and basally ovoid (Figs. 1C, 5D, F; fig. 5 Wunderlich 2011), and the mesal 
bristle of the embolic apophysis protmding from beneath the conductor (Figs. 1C, 4D-E; figs. 3-5 
Wunderlich 2011). The former is also shared with Wendilgarda (fig. 198 Coddington 1986). 
Females of Coddingtonia have lateral oblique pockets on the epigynmn (Figs. 6D, 7D; fig. HE 
Miller et al. 2009), different to the lateral round sclerotized pits found in Alaria and Theridioso- 
matinae (fig. 2B Zhao and Liu 2012; figs. 151, 173, 213 Coddington 1986). 
Diagnosis. — Coddingtonia and Tagalogonia gen. nov. can be distinguished from other gen¬ 
era of Theridiosomatidae by the opisthosoma sparsely covered with long and strong setae, the well- 
developed disk shaped conductor, the long and slender cymbium with a ventral groove where the 
conductor fits and the long and thin copulatory ducts (Figs. lA-D, 2A-F, 3A-F, 4A-F, 5A-F, 
6A-F, 7A-F; figs. 8B, 1 lE-F Miller et al., 2009; fig. 3 Wunderlich, 2011). Males of Coddingtonia 
can be distinguished by having the mesal bristle of the embolic apophysis protruding from beneath 
the conductor (Figs. 1C, 5D-E; fig. 3 Wunderlich 2011), the median apophysis elongated, ovoid 
and distally acute (Figs. IC, 5D, F; fig. 5 Wunderlich 2011), and the striated tegulum (Fig. 1C; fig. 
4 Wunderlich 2011), while those of Tagalogonia gen. nov. have the embolus and its apophysis cov¬ 
ered by the conductor (Figs. lA, 2E, 4F), the median apophysis reduced and square (Figs. 1 A, 2F, 
4F), and lack denticules on the tegidum (Fig. lA). Females of Coddingtonia resemble those of 
Tagalogonia gen. nov. by the separated spennathecae, the epigynum with transverse ridges and a 
deep central transversal pit (Figs. IB, C, 3D-F, 6D-F, 7D-F; figs. IIE-F Miller et al. 2009; figs. 
6-7 Wunderlich 2011). However, Coddingtonia females can be distinguished by the pattern of the 
copulatory ducts that describes a posterior-lateral loop, followed by a dorsal loop, and finally a pos¬ 
terior-median loop that coils more than four times before inserting on the round or oval spermath- 
ecae mesally (Figs. ID, 6E, 7E; fig. IIF Miller et al. 2009), by the lateral wings of the copulatory 
bursae well developed, like round, swollen sacks, with dorso-median gland ductiles (Figs. ID, 
6E-F, 7E-F; fig. IIF Miller et al. 2009; fig. 7 Wunderlich 2011), and by the central pit on the ante¬ 
rior third of the epigynum above the spennathecae (Figs. ID, 6D, 7D; fig. HE Miller et al. 2009). 
In Tagalogonia gen. nov. the copulatory ducts describe a lateral loop, followed by a posterior-medi¬ 
an loop, and finally an anterior-median loop that inserts on the renifonn spennathecae mesally 
(Figs. IB, 3D, F), the lateral wings are soft, not swollen, without gland ductiles (Figs. IB, 3D, F), 
and the central pit of the epigynum is in the middle line under the spermathecae (Figs. IB, 3D-E). 
