432 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
(Figs. 5 A-C). Ozopore fonnula as other Polydesmida: rings 5*, 7^^ 9^^, 10^'^ 12^^, 13*, 15^^19^^. 
Ozopores simple, sub-circular, opening dorsally on paranoia, posterior to transverse median line of 
metazonite (Figs. 6 A-B). Sternites as wide as long, with disperse brush-tipped twisted setae. Ante¬ 
rior tracheal spiracles opening anteriorly, and posterior tracheal spiracles opening posteriorly. Tra¬ 
cheal spiracles clavate, extending dorsally as long as the diameter of leg socket (Fig. 5 D ). Spirac- 
ular filter coral-like (Figs. 5 D-E). Pre-anal ring: Epiproct well developed, wider than long, semi¬ 
circular and flat; with seven marginal setae (Fig. 1 D), and four spinnerets. Paraprocts with two 
elongated and filiform setae each, which crosses median edge fonned by paraprocts (Fig. 1 D). 
Dorsal edge of hypoproct truncated with two elongated filiform setae (Fig. 1 D). Legs: Legs cov¬ 
ered with brush-tipped t^^dsted setae, especially abundant on dorsal suiTace (Figs. 6 A, D, G-H). 
First and second leg prefemora with strong and filifonn setae, which are as long as width of seg¬ 
ments (Figs. 6 A, D). Length of first leg coxa and of prefemur to tarsus, y4 times longer than other 
legs (Fig. 6 A). Surface of first leg coxacoarse with dispersed small peaks (Fig. 6 C). Ventral sur¬ 
faces of first and second coxae, pre-, femora and postfemora covered with long filifonn and brush- 
tipped twisted setae; tibiae to tarsi with short filifoim setae (Figs. 6 A, D). Tarsal setae short (Figs. 
6 A, D, G-H). Coxa of second leg bears a single smooth gonopore (Figs. 6 D-F). Legs of succeed¬ 
ing segments covered with short, brush-tipped twisted setae (Figs. 6 G-H). Prefemur with single 
long proximal filiform setae (Fig. 6 G). Go nopod: Gonocoxae massive, twice as long as wide, 
slightly joined along their median edges, outside of segment trapezoidal aperture and with elon¬ 
gated bmsh-tipped twisted setae anteriorly and laterally (Figs. 7 A-D, 8 A-D). Prefemur with dense 
filifonn, strong setae anteriorly, and with thin setae close to cannula (Figs. 7 B, D, 8 B, D). 
Telopodite pivoting on gonocoxae, twisted clockwise; with two branches (Figs. 7 A-D, 8 A-D). 
Solenomere extended anteriorly and curved posteriovenrially, recurving dorsally; distally acute; tip 
pointing mesodorsally (Figs. 7 A, C, 8 A, C; “A” process in Hoffman, 2001). Tibiotarsus base wide, 
distally acute, pointing dorsally, with four bumps on posterior edge (Figs. 7 C-D, 8 C-D; “D” 
process in Hoffman, 2001). Epigyme: Epigyne rectangular, with a sculptured edge, not rising above 
the ventral surface of ring 3 (Fig. 9 A). The center of the epygine extends ventrally on a rounded- 
triangular prong (Fig. 9 A). 
Remarks. — Eggs spherical, surface finely wrinkled (Figs. 9 B). 
Variation.— Males (n = 4). Females (n = 3): Body length/width between 6.5-7.3. Labium 
with two or three teeth. 
Distribution. — Philippines: Luzon Island: Mount Makiling, Los Banos (Laguna Province), 
and Manila. Wang (1951) described several males and females of /. margaritiferus from Mindanao 
Island, Philippines. However, as was mentioned in the Systematics section, what Wang (1951) re¬ 
described and illustrated as P. philippinorum instead has the diagnostic characters of /. margan- 
tiferus, i.e., a solenomere extended anteriorly, curved postero-ventrally and recurved dorsally, and 
a tibiotarsus pointing dorsally with four bumps on the posterior edge (toothed) (Wang 1951, fig. 
71). In contrast, what Wang re-described as P margaritiferus has a solenomere with a single curve 
(not secondary curvature), and the tibiotarsus lacks teeth (Wang 1951, fig. 73). Thus, Wang’s 
P. margaritifems specimens should not be considered in the distribution of /. margaritiferus. Suma¬ 
tra: Karsch (1881) described the gonopod Ifom a male from Sumatra, which we did not examine. 
As Jeekel (2007) suggested, it seems doubtflil that the specimen is a I. margaritiferus. 
Acknowledgments 
This research was facilitated by a generous gift to the CAS from Will and Margaret Hearst sup¬ 
porting the 2011 Filipino-American Hearst Biodiversity expedition to Luzon. We thank the other 
