448 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
This morphospecies, represented by a single female in our sample, may be conspecific with die preceding 
species, represented by a single male; but we separate the two here because they have distinctly different 
elytral topographic, macrosculptural and microsculptural features (compare Figs. 24 and 25), and one 
species having so many gender-specific differences would be unusual. 
Chlaenius (Lissauchenius) sp. (Fig. 26), 1 specimen; PH0051; collected at mercury vapor light in forest 
Harpalini 
Stenolophus {Egadroma) sp. (Fig. 27), 2 specimens; PH0006, PH0015; collected by mini-Winkler extraction 
from concentrated leaf litter and at ultraviolet light, both in forest 
Acupalpus sp. (Fig. 28), 1 specimen; PH0004; collected in pitfall traps in forest. 
Coleolissus sp. A (Fig, 29), 1 specimen; PH0051; collected at mercury vapor light in forest 
Coleolissus sp. B (Fig. 30), 1 specimen; PH0043; collected at mercury vapor light in grassland 
Coleolissus sp. C (Fig. 31), 25 specimens; PH0009, PHOOlO, PHOOl 1, PH0015; collected by hand in daytime, 
in pitfall traps and by mini-Winkler extraction from concentrated leaf litter, all in forest 
Coleolissus sp. D (Fig. 32), 2 specimens; PH0006, PH0032; collected at ultraviolet and mercury vapor hghts 
in forest 
Trichotichnis sp. (Fig. 33), 1 specimen; PH0025; collected by mini-Winkler extraction from concentrated for¬ 
est leaf litter 
Orthogoniini 
Orthogonius sp. A (Fig. 34), 1 specimen; PH0050; collecting by hand in daytime in forest. 
Orthogonius sp. B (Fig. 35), 1 specimen; PH0020; collected by hand at night in forest 
Platynini 
Agonum muelleri Herbst (Fig. 36), 1 specimen; PH0027; collected in yellow pan traps in forest 
This is a Palaearctic species, which has been introduced into North America on both coasts and continues 
to spread its range on that continent. To the best of our knowledge, it has not been recorded from anywhere 
in eastern Asia, so its occurrence in our samples is quite unexpected. If our identification is conect, then 
this must represent an introduction into die Philippines, perhaps even fr'om North America; and it may or 
may not actually be established in the islands. 
Altagonum sp. A (Fig. 37), 4 specimens; PH0021, PH0034, PH0037; collected by beating vegetation and at 
ulfraviolet light, all in forest 
Altagonum sp. B (Fig. 38), 2 specimens; PH0020, NCP_3; collected by hand in forest in daytime and at night 
Altagonum sp. C (Fig. 39), 8 specimens; PH0021, PH0023, PH0034; collected by hand at night, by beating 
vegetation, and at ultraviolet light, all in forest 
Altagonum sp. D (Fig. 40), 1 specimen; PH0061; collected by mini-Winkler extraction of concentrated for¬ 
est leaf litter 
Arhytinus minimus Jedlicka (Fig. 41), 2 specimens; PH0015; collected by mini-Winkler extraction of con¬ 
centrated forest leaf litter 
Colpodes sp. A (Fig. 42), 2 specimens; PH0021; collected at ultraviolet light in forest 
Colpodes sp. B (Fig. 43), 1 specimen; PH0021; collected at ultraviolet light in forest 
Gastragonum sp. (Fig. 44), 19 specimens; PH0024, PH0055; collecting by hand in cryptic habitats in forest 
To the best of olu knowledge, this genus has not been recorded previously outside of New Guinea. 
Tliis is one of Darlington’s (1952) “genera of convenience”, based mainly on dorsal chaetotaxic features, 
so we assign this morphospecies to Gastragonum very tentatively and only because that’s where it keys 
out in his key to genera. 
Notagonum sp. A (Fig. 45), 10 specimens; PH0029, PH0054, PH0062, PH0063; collected by hand at night, 
in pitfall traps, and at ultraviolet light, all in forest 
Notagonum sp. B (Fig. 46), 2 specimen; PHOOOl, PH0004; collected in daytime by hand and in pitfall traps, 
all in forest 
Notagonum sp. C (Fig. 47), 2 specimens; PHOOl8; collected in daytime by hand in forest 
