450 
THE CORAL TRIANGLE: HEARST BIODIVERSITY EXPEDITION 
Dryptini 
Dendrocellus coelestinus (Klug) (Fig. 67), 1 specimen; PH0040; collected by hand sorting of litter sifted 
from grassland and degraded forest 
This represents a first record for this species in the Phillippine Islands (Liang and Kavanaugh 2007). 
Brachinini 
Brachinus sp. (Fig, 68), 2 specimens; MHW_2; collected by hand at night in forest 
Both published (Maddison et al 1999) and unpublished molecular data (D. R. Maddison, personal commu¬ 
nication) strongly support the Brachininae as sister group to the huge subfamily Harpalinae. Hence, we 
place the brachinine species here in our list rather than with the Paussinae as done by Lorenz (2005). 
Acknowledgements 
First of all, we must thank Will and Margaret Hearst for their generosity in funding the Aead- 
emy’s 2011 Philippine Expedition. We gratefully acknowledge the many colleagues who collected 
the specimens on which this report is based. Among these are Natalia Chousou Polydouri, Charles 
Griswold, Vanessa Knutson, Matt Lewm and Hannah Wood of the California Academy of Sciences 
team, and O. Eusebio, V. San Juan, S. Yap, M. Yngente of the Philippine contingent. We also thank 
Alma Saucedo, who prepared the carabid specimens for this project, and Martin Baehr (Platynini, 
Pentagonicini andLebiini), Terry Erwin (Tachyina and Brachinini), David Maddison (Bembidiini), 
and Kipling Will (Pterostichmi) for their help with identifications. 
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