Occasional Papers Series of the California Academy of Sciences 
Alan E. Leviton, Editor 
Gary C. Williams, Associate Editor 
Michael T. Ghiselin, Associate Editor 
Peter Roopnarine, Associate Editor 
Michele L. Aldrich, Consulting Editor 
Copyright © 2012 by the California Academy of Sciences, 55 Music Concourse Drive, San Francisco, Califor¬ 
nia 94118. Date of publication: 1 June 2012. 
All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, 
electronic or mechanical, including photocopying, recording, or any information storage or retrieval system, 
without permission in writing from the publisher. 
ISSN 0068-5461 
Printed in the United States of America 
Allen Press, Lawrence, Kansas 66044 
