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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 8 
but this is not currently supported by voucher 
specimens. 
Libyan Records (Map 8): CYRENAICA: 
Kufrah : 581: BMNH 1975.1228. 582: Sinda- 
co and Jeremcenko 2008; Trape et al. 2012. 
583: MZUF 721. 584: Scortecci 1935c; Zavat- 
tari 1937. 586: MZUF 722; Scortecci 1935c. 
587: MZUF 720; Scortecci 1935b,c, 1939. 
Comments.— This recently described 
species was previously included within 
Pseudotrapelus sinaitus (Heyden, 1827), a 
species now restricted to the Sinai Peninsula, 
parts of Jordan, southern Syria, western Iraq, 
and northwestern Saudi Arabia (Tamar et al. 
2016a). The original authors of P. chlodnickii 
(Melnikov et al. 2015) provided only a few 
localities for the taxon, all in Sudan. As noted 
by Tamar et al. (2016a), the piecemeal reevalu¬ 
ation of the Pseudotrapelus sinaitus complex 
by Melnikov and colleagues (Melnikov et al. 2012, 2013a, 2013b, 2014, 2015; Melnikov and Pier¬ 
son 2012; Melnikov and Melnikova 2013; Melnikova et al. 2015), based on small samples form 
restricted localities, left the taxonomic status of many populations of Pseudotrapelus unresolved. 
Tamar et al. (2016a) sampled sufficiently broadly to confirm that a specimen from southwestern 
Egypt, close to the Kufrah records, was conspecific with near-topotypical P. chlodnickii. 
Le Berre (1989) did not include Libya in his map of the distribution of Agama sinaita, but 
explicitly mentioned Scortecci’s (1939) record from Jabal Al Uwaynat. This locality straddles the 
Libyan, Sudanese and Egyptian borders and historical records from Jabal Al Uwaynat and Ayn 
Murr have been variously interpreted by different authors as being Libyan (e.g., Kramer and 
Schnurrenberger 1963) or Sudanese (Baha El Din 2006a). We here treat the record as Libyan. 
IUCN Threat Status.— Not evaluated, but anticipated to be Least Concern. 
Trapelus mutabilis (Merrem, 1820) 
Trapelus mutabilis mutabilis (Merrem, 1820:50) (Fig. 12) 
1820 Agama mutabilis Merrem, Versuch eines Systems der Amphibien — Tentamen Systematis Arnphi- 
biomm. Johann Christian Krieger, Marburg, Germany. XV + 191 pp. [German], XV + 191 pp. [Latin], 1 pi. 
Neotype.— ZFMK 64395 designated by Wagner et al. (2011), “Egypt, 10 km northwest of Cairo.” Orig¬ 
inal type locality “Egypto’7”in Aegypto” (Merrem, 1820). Merrem (1820) based his description on specimens 
figured by Geoffroy Saint-Hilaire (1809, pi. 5, figs. 3-4) in the “Description de l’Egypte”. According to Wag¬ 
ner et al. (2011) there is no corresponding type in MNHN, where specimens figured in the “Description de 
VEgypte ” would be expected to be deposited, nor in the University of Marburg, where Merrem had been 
based. Likewise there are no possible types in BMNH, where Baha El Din (2006) suggested the type was 
housed. 
Trapelus mutabilis poppeki Wagner, Melville, Wilms and Schmitz 2011:896, fig. 7 
2011 Trapelus mutabilis poppeki Wagner, Melville, Wilms and Schmitz, Opening a box of cryptic taxa - 
the first review of the North African desert lizards in the Trapelus mutabilis Merrem, 1820 complex (Squa- 
mata: Agamidae) with descriptions of new taxa. Zoological Journal of the Linnaean Society, 163:884-912. 
Map 8. Distribution of Pseudotrapelus chlodnickii in 
Libya. 
