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Figure 12. Trapelus mutabilis mutabilis from 5 km south of Gadamis, Nalut, Tripolitania, Libya, 2008. Photo © Rober¬ 
to Sindaco. 
Holotype.— ZFMK 63678, “Libya: east of Tarbu, 24.48.27N, 16.19.11E, 490m a.s.l.” 
Agama mutabilis [part], Le Berre 1989:116. 
Trapelus mutabilis [part], Schleich, Kastle and Kabisch 1996:291 
Trapelus mutabilis [part], Sindaco and Jeremcenko 2008:162. 
Complexe Trapelus mutabilis, Trape, Trape, and Chirio 2012:162. 
Distribution.— Following the taxonomic conclusions of Wagner et al. (2011), this species as 
a whole occurs from Algeria east through Libya to the Nile ( T. m. mutabilis ), and eastwards to Iraq, 
Kuwait and the western shore of the Persian Gulf (T. m. pallidus ). Trapelus m. poppeki is known 
only with certainty from two disjunct localities in western Libya. 
Libyan Records (Map 9): ( Trapelus mutabilis mutabilis ): TRIPOLITANIA: Zawiyah : 11: 
BMNH 1965.1186-87. 22: MCSN 3029. Jafara : 28: Andreucci 1913. Tripoli : 36: BMNH 
1954.1.5.63-65. 39: FMNH 82950. 44: SMNS 438. 45: MCSN 4231; MZUT R313; ZMB 15276- 
80, 15296, 15316-17, 54582, 54585-86, 54591-97; Condorelli-Francaviglia 1896; Werner 1909; 
Andreucci 1913; Ghigi 1913; Zavattari 1934. 46: Sayers 1964. Murqub : 68: Peters 1880, 1881. 
Misratah : 80: MSNG 52507; Boulenger 1914; Zavattari 1934. 88: RMNH RENA 13915. Nalut : 
101: Sindaco, pers. obs. 4/30/2008. 103: Sindaco, pers. obs. 4/30/2008. 104: BMNH 1965.1183. 
105: AIC no number provided; MNHN 2004.0085; Ibrahim and Ineich 2005. 109: BMNH 
1965.1185. 113: Sindaco, pers. obs. 5/2/2008. 114: BMNH 1954.1.5.66. 116: Sindaco, pers. obs. 
5/2/2008. 130: Sindaco, pers. obs. 4/1/2008. Jabal al Gharbi : 137: ZSM 116/1994/1, 116/1994/2. 
140: BMNH 1965.1184. 143: NHMC 80.3.90.20. 154: MCSN 4230; MZUT R336; Andreucci 
1913; Zavattari 1934. 160: BMNH 1954.1.5.67. 163: Brito et al. 2008. Sirte : 174: MCSN 1947; 
Peters 1880, 1881. 175: ZMB 37010, 54429-30. 199: MCSN 1832; MCSN 4229. “Tripolitania 
settentrionale”: Zavattari 1937. “Ras Wagis”: BMNH 1901.10.28.4. FEZZAN: Jufra : 213: 
BMNH 1954.1.5.69. 216: MCSN 1950. 218: MCSN 1778. 226: MCSN 1780, 4226; Scortecci 
1935b. 228: BMNH 1954.1.5.68. Ghat : 234: Essghaier et al. 2015. 252: MCSN 1843. 253: MCSN 
1839. Wadi alHayaa : 269: Essghaier et al. 2015. Sabha : 286: BMNH 1954.1.5.72. 291: NMBA- 
