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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 6 
Table 1. Comparative morphology of Rostanga species. 
Species 
Rhinophoral 
lamellae 
Jaw 
rodlets 
Inner lateral 
Middle lateral 
Outer lateral 
Vestibular 
gland 
alisae 
vertical 
present 
denticulate 
smooth 
multifid 
absent 
aliusrubens 
vertical 
present 
denticulate 
smooth, inners 
multifid 
absent 
ankyra 
transverse 
absent 
bifid 
bifid 
bifid 
absent 
anthelia 
vertical 
present 
denticulate 
smooth 
multifid 
? 
arbutus 
transverse 
present 
denticulate 
denticulate 
multifid 
absent 
aureomala 
transverse 
absent 
bifid also rachidian 
present 
bifid 
bifid or trifid 
absent 
crawfordi 
vertical 
reduced 
denticlulate or smooth 
smooth 
multifid 
absent 
bassia 
vertical 
present 
denticulate 
no denticles 
multifid 
absent 
bifurcata 
transverse 
present 
denticulate 
denticulate 
bifid 
absent 
byga 
vertical 
reduced 
denticulate 
smooth 
multifid 
absent 
calumus 
vertical 
present 
denticulate 
no denticles 
multifid 
absent 
crocea 
transverse 
absent 
bifid 
bifid 
bifid 
? 
dentacus 
transverse 
reduced 
denticulate 
no denticles 
undivided or bifid 
absent 
elandsia 
transverse 
absent 
bifid 
bifid 
bifid 
absent 
ghiselini 
transverse 
reduced 
denticulate 
bifid 
bifid 
present 
lutescens 
transverse 
present 
denticulate 
smooth 
trifiid 
present 
muscula 
vertical 
reduced 
denticulate 
no denticles 
multifid 
present 
orientalis 
vertical 
present 
denticulate 
no denticles 
multifid 
absent 
phepha 
transverse 
reduced 
bifid 
smooth 
bifid 
absent 
pulchra 
vertical 
reduced 
denticulate 
smooth 
mutifid 
absent 
risbeci 
transverse 
reduced 
denticulate 
smooth 
bifid 
absent 
rosi 
transverse 
absent 
smooth 
smooth 
smooth 
present 
rubra 
vertical 
present 
denticulate 
bifid 
bifid 
absent 
setidens 
transverse 
absent 
smooth 
smooth 
bifid 
absent 
verse lamellae, whereas those of R. pulchra are broad with vertical lamellae. There are 6-7 bipin- 
nate gill branches in R. ghiselini and 8-12 unipinnate branches in R. pulchra. Internally, the radu- 
lar morphology of the two species differs. The inn ermost radula tooth is denticulate in both species 
but in R. ghiselini the tooth bears 8-21 denticles whereas in R. pulchra there are 7-9 denticles (Fig. 
5E, F). In R. ghiselini all of the hamate teeth are strongly bifid whereas only the second tooth is 
bifid in R. pulchra. The 67 elongate outer teeth of R. ghiselini have bifid apices whereas the 25 
elongate teeth have 3-6 fine apices in R. pulchra (Fig. 5H). The reproductive system is similar in 
both species with the exception that the vagina of R. pulchra is more elongate. 
Valdes and Gosliner (2001) included Boreodoris Odhner, 1939 as a synonym of Rostanga, not¬ 
ing that the absence of jaws and a denticulate inner lateral tooth was not sufficient to warrant gener¬ 
ic separation. Subsequently, Garavoy, Valdes and Gosliner (2001) described three new South 
African species of Rostanga that all had a bifid inner lateral tooth. Two of these species, R. aureo- 
mala and R. elandsia, lacked jaws, whereas R. phepha had distinctive jaw rodlets. Martynov 
(2003), stating that Boreodoris lacking jaws and lacking an innermost lateral tooth with denticulate 
flange, is sufficiently distinct from Rostanga and should be considered as a distinct genus. Based 
