200 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 8 
species, the second as a synonym of the third, and considered another member of the group, 
T. somalicus Parker, 1942, as a full species as well. The nominate form occurs in Libya and the 
eastern portions of the species range as far west as Tunisia. The Libyan distribution of this species 
has been variously interpreted. Le Berre (1989) showed a near coastal distribution and an isolated 
area corresponding to northern A1 Kufrah, along the Egyptian border. Schleich et al. (1996) con¬ 
sidered the species to be limited in Libya to Cyrenaica. We concur with Sindaco and Jeremcenko 
(2008) that the species occurs throughout most of Libya. Note that Ibrahim (2008a) listed the same 
specimen number (ZCT 2005.16) for both localities 386 and 387. 
IUCN Threat Status.— Least Concern. 
Family Phyllodactylidae 
Ptyodactylus togoensis Tornier, 1901:68 (Fig. 19) 
1901 Ptyodactylus hasselquisti var. togoensis Tomier, Die Crocodile, Schilkroten und Eidechsen in Togo. 
Archiv fur Naturgeschichte 1901 Beiheft:65-88. 
Syntypes. — ZMB 16312, 16622, “Mangu” [= Mango, formerly Sansanne-Mango,Togo]. 
Ptyodactylus hasselquistii [part], Le Berre 1989:158. 
Ptyodactylus ragazzii [part], Schleich, Kastle, and Kabisch 1996:244. 
Ptyodactylus ragazzii [part], Sindaco and Jeremcenko 2008:124. 
Ptyodactylus ragazzii [part], Trape, Trape, and Chirio 2012:266. 
Distribution.— Mauritania, Mali and Niger through southern Algeria, southwestern Libya, 
and northern Chad. South across the Sahel from Burkina Faso to Togo and Nigeria, northern 
Cameroon and extreme southwestern Chad. 
Libyan Records (Map 20): FEZZAN: Ghat : 237: Sindaco, pers. obs. 4/20/2008. 241: Sinda¬ 
co, pers. obs. 4/21/2008. 240: Sindaco, pers. obs. 4/21/2008. 249: CUP R 080-81; MCSN 1971- 
Figure 19. Ptyodactylus togoensis from Awiss area, Tadrart Akakus, Ghat, Fezzan, Libya, 2008. Photo © Roberto Sin¬ 
daco. 
