BAUER, DEBOER, & TAYLOR: ATLAS OF THE REPTILES OF LIBYA 
201 
72, 1975-76 ; ZCT 2006.72; Zavatari 1934, 
1937; Scortecci 1937a; Loveridge 1947; Fryn- 
ta et al. 2000; Ibrahim 2008a. 251: CUP R Oil, 
072-79, 103; Frynta et al. 2000. 257: Sindaco, 
pers. obs. 4/21/2008. 261: Sindaco, pers. obs. 
4/22/2008. 263: CUP R 071; Frynta et al. 2000. 
“Zone of Ghat”: Scortecci 1934b. Murzuo : 
298: Essghaier et al. 2015. 303: Zavattari 1937. 
313: AIC 2006.1573, ZCT 2006.47-48; 
Ibrahim 2008a. 316: NMBA-REPT 15301-03; 
ZCT 2006.63-64, 69; Zavattari 1934; Ibrahim 
2008a. 
Comments. — The distribution of Ptyo- 
dactylus ragazzii according to Sindaco and 
Jeremcenko (2008) and subsequently modified 
(Riva and Padial 2008; Perera and Harris 2010) 
extends from Mauritania to the Horn of Africa. 
Metallinou et al. (2015) demonstrated that 
western populations of P. “ragazzii ” as far east as the Tassili, Ahaggar (Algeria), and Air Moun¬ 
tains (Niger) were not sister to true P. ragazzii from Djibouti and Ethiopia and resurrected the name 
P. togoensis from synonymy for these western forms, which are themselves geographically and 
genetically fragmented (Froufe et al. 2013). However, the sampling gap of Metallinou et al. (2015) 
included all of Libya. Thus, genetic information is lacking regarding the identity of Libyan speci¬ 
mens of Ptyodactylus. Based on the relative proximity of southwestern Libyan records to P. togoen¬ 
sis in the Tassili and elsewhere in southern Algeria, we hypothesize that Libyan material can be 
referred to this species, although this is subject to genetic confirmation. 
IUCN Threat Status. — Not evaluated, but anticipated to be Least Concern. 
Tarentola annularis (I. Geoffroy Saint-Hilaire, 1827:130) 
1827 Gecko annularis Geoffroy Saint-Hilaire, Description des reptiles qui se trouvent en Egypte, pp. 
121-160 in M. J.-C. L. de Savigny, (ed.), Description de FEgypte, ou Recueil des Observations et des 
Recherches qui ont ete Faites en Egypte Pendant l’Expedition de l’Armee Frangaise. Histoire Naturelle. Tome 
premier. Partie premier. Imprimerie Imperiale, Paris, France. 
Lectotype: — MNHN 6697a designated by Joger (1984), “Egypte” by implication. 
Tarentola annularis, Le Berre 1989:168. 
Tarentola ( Sahelogecko ) annularis, Schleich, Kastle, and Kabisch 1996:266. 
Tarentola annularis, Sindaco and Jeremcenko 2008:133. 
Tarentola annularis, Trape, Trape, and Chirio 2012:268. 
Distribution. — North Africa from Western Sahara, Mauritania and Senegal through Mali, 
Niger, Burkina Faso, northern Cameroon, Central African Republic, and Sudan to Ethiopia, Eritrea 
and Somalia but absent from most of Morocco, Algeria and Libya (Sindaco and Jeremcenko 2008). 
Recently introduced into Israel (Jamison et al. 2017). In Libya they are found scattered across the 
southern portions of the country. 
Libyan Records (Map 21): FEZZAN: Murzuo : 316: NHMW 17969. CYRENAICA: 
Kufrah : 574: MZUF 693; Scortecci 1935c; Loveridge 1947. 575: Loveridge 1947; Joger 1984. 
584: MZUF 692; Scortecci 1935c; Zavattari 1937; Loveridge 1947; Joger 1984. 
Map 20. Distribution of Ptyodactylus togoensis in Libya. 
