BAUER, DEBOER, & TAYLOR: ATLAS OF THE REPTILES OF LIBYA 
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Zavattari 1937. “Sirtica”: Zavattari 1937. FEZZAN: Wadi al Shatii : 200: Brito et al. 2008. 210: 
NMBA-REPT 15470; ZMB 37603. 211: Zavattari 1937. Jufra : 213: BMNH 1954.1.6.3; ZSM 
165/1979, 175/1979; Salvador 1982. 216: ZMB 18008; ZSM 134/1983; Peters 1880, 1881; Wern¬ 
er 1909; Ghigi 1913; Zavattari 1934. 217: Zavattari 1937. 218: MCSN 2062, 2073; ZSM 44/1981. 
221: MCSN 2074; Scortecci 1935b. Ghat : 249: MCSN 2069,2077,2168. 253: MCSN 2067,2113; 
Scortecci 1937a. Wadi al Hayaa : 265: MCCI R1452(l); Brito et al. 2008; Sindaco, pers. obs. 
4/24/2008; Tamar et al. 2016b. 267: Sindaco, pers. obs. 4/24/2008. 268: Brito et al. 2008. 269: 
Zavatarri 1934. 270: Sindaco, pers. obs. 4/24/2008. 272: MCSN 2061. 274: CUP R 025; Frynta et 
al. 2000. Sabha : 281: MCSN 2070. 285: MCSN 2078; MSNG 50658; NMBA-REPT 15428-31; 
Kramer and Schnurrenberger 1963; Schnurrenberger 1963. 291: NMBA-REPT 15310-17. 
Murzuo : 298: AIC 2006.1572; BMNH 1954.1.5.9; MCSN 2076, 2165; ZSM 161/1979; Salvador 
1982; Ibrahim 2008; Werner and Ibrahim 2012. 303: BMNH 1956.1.1.7; MCSN 2060; ZCT 
2005.04; Salvador 1982; Ibrahim 2008a. 311: ZCT 2006.33, 2006.41, 2006.66; Ibrahim 2008a. 
312: MCSN 2083. 316: ZSM 135/1983. 318: Kramer and Schnurrenberger 1963; Schnurrenberger 
1963. “Fezzan”: BMNH 1954.1.5.91-97; Salvador 1982. CYRENAICA: Benghazi : 342: MCSN 
2173; MZUT R2278; USNM 146795; Salvador 1982. 357: BMNH 1954.1.5.79-86; MTKD 
17224-27; MZUF 769; NHMW 8216; SMF 34696; von Martens 1883; Werner 1909; Ghigi 1913; 
Calabresi 1923; Umani 1923; Zavattari 1929,1930, 1934; Salvador 1982. 361: BMNH 1987.1029. 
363: BMNH 1985.1191-92. 365: SMF 34588. 376: BMNH 1954.1.5.87-90; MCSN 2469; Sal¬ 
vador 1982. Marj : 385: MCSN 1943; Zavattari 1937. 389: SMF 34587; Calabresi 1923; Zavattari 
1929, 1930, 1934. Jabal alAkhdar : 410: FMNH 82940. 414: FMNH 82935, 82939, 82967-68. 
415: FMNH 214941-44; Resetar 1981. 417: Zavattari 1922, 1929, 1934; Calabresi 1923. 452: 
Zavattari 1929, 1930. 457: KNP 1981/149, 204, 277, 292, 298-299, 305-308, 311, 318, 324, 329, 
339-341, 343-344, 347-349, 353, 357, 394, 396-397, 406, 409; Schleich 1987; 457f: Schleich 
1987. 4571: Schleich 1987. 4571: Schleich 1987. Darnah : 484: MCSN 2364; MZUF 661, MZUT 
R2276; Calabresi 1923; Zavattari 1929, 1930, 1934. Butnan : 507: BMNH 1965.1251-53; Sal¬ 
vador 1982. Al Wahat : 529: MSNG 31604*; MCSN 2017, 2064, 2081; ZSM 138/1983; Vin- 
ciguerra 1931; Zavattari 1934; Scortecci 1935b. 533: MSNG 31604*. 537: SMF 55755. 540: 
BMNH 1965.1254; Salvador 1982. 544: MSNG 31604*; Vinciguerra 1931; Zavattari 1934. 553: 
MCSN 2172; ZSM 137/1983. 557: BMNH 1909.7.28.28-29; Salvador 1982. 558: NMP 34932; 
Moravec 1995. 564: Zavattari 1929, 1930, 1934,1937. “Cyrenaica”: ZMB 10502. “Kouf Nation¬ 
al Park, Ras Latin”: ZSM 136/1983. LIBYA: NHMW 31123; NMP 34932. 
Comments. — The complicated dating of the natural history portions of Description de 
I’Egypte has been elucidated by Sherbom (1897) and Tollitt (1986). The subspecies present in 
Libya and most of the North African range is A. b. asper. The A. boskianus species complex has 
long been believed to be paraphyletic (Harris and Arnold, 2000). The nominate form occupies the 
region of the Nile Delta and coastal north Sinai. Acanthodactylus b. euphraticus Boulenger, 1919 
is in the region of Iraq. Trape and Trape (in Trape et al. 2012) named A. b. khattensis from west¬ 
ern Mauritania and Trape, Chirio, and Geniez (in Trape et al. (2012) erected A. b. nigeriensis for 
material from Niger. The subspecific division of A. boskianus has been considered questionable 
(Sindaco and Jeremcenko 2008), but Wemer and Ibrahim (2012) presented morphological evidence 
for the distinctness of A. b. asper and A. b. boskianus. Tamar et al. (2016b) recovered a “ boskianus 
group” within which A. boskianus asper was made paraphyletic by A. schreiberi schreiberi + A. s. 
ataturi, A. s. syriacus, A. b. nigeriensis, and A. b. khattensis. Divergences within A. b. asper are 
relatively shallow. The sole Libyan sample in the study of Tamar et al. (2016b), from the southwest 
of the country, is in a clade including samples from Mauritania, Niger, Western Sahara, Morocco, 
Algeria, Tunisia, Egypt (west of the Suez Canal) and Sudan. This clade represents true A. b. asper 
