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Series 4, Volume 64, No. 8 
and its inclusion of both A. b. nigeriensis and A. b. khattensis suggests that these two taxa should 
be synonymized with A. b. asper (Tamar et al. 2016b). Material from Sinai, Israel, Jordan and 
Lebanon (including some specimens identified as A. s. syriacus ), Yemen and southern Oman also 
form a relatively shallow clade and although labeled as A. b. asper in Tamar et al.’s (2016b) tree, 
correspond to the traditional concept of A. b. boskianus. Other clades in the A. boskianus group 
include: 1) specimens from the U.A.E., northern Oman, and the Jordanian panhandle, 
2) A. schreiberi schreiberi + A. s. ataturi from Cyprus and the Cilician region of Turkey, respec¬ 
tively, and 3) Syrian A. boskianus. 
IUCN Threat Status. — Not assessed, but assumed to be Least Concern. 
Acanthodactylus dumerilii (Milne-Edwards, 1829:85, pi. VII, fig. 9) 
1829 Lacerta Dumerilii Milne-Edwards, Recherches zoologiques pour servir a l’histoire des lezards, 
extraites d’une monographic de ce genre. Annales des Sciences Naturelles, Paris, ser. 1, 16: 50-89, pis. 
V-VIII. 
Lectotype. — MNHN 2759 fide Brygoo (1988), “Senegal” designated by Crochet et al. (2003). 
1838 Scapteira inornata Gray, Catalogue of the slender-tongued saurians, with descriptions of many new 
genera and species. Part 1. An nals and Magazine of Natural History, series 1, 1:274-283 [281]. 
Lectotype. — BMNH 1946.9.3.76, “Tripoli” [Libya] designated by Crochet et al. (2003). 
1885 Acanthodactylus scutellatus var. exiguus Lataste, Les acanthodactyles de Barbarie et les autres 
especes du genre. Description d’une nouvelle espece, du pays des (fomalis ( Acanthodactylus Vaillanti ). Annali 
del Museo Civico di Storia Naturale di Genova, ser. 2, 2:476-516 [493]. 
Lectotype. — BMNH 1920.1.20.1349, designated by Crochet et al. (2003), “en Algerie, a Biskra, 
Mraier, Tougourt, Bled Ahmar, Hadjira, N’Gou 9 a, Tilremt, Laghouat, Am-el-Hel et Bou-Saada [...] en 
Tunisie, a l’oued el Kreil (entre le plateau de Haouaia et Ghumra 9 en), Kebili (dans le Nefzaoua), Tozeur, et 
Nefta (dans le Djerid),” restricted to “Hadjira, Algeria” by lectotype designation. 
Acanthodactylus scutellatus [part], Le Berre 1989:190. 
Acanthodactylus dumerili, Sindaco and Jeremcenko 2008:220. 
Complexe Acanthodactylus dumerili, Trape, Trape, and Chirio 2012:304. 
Distribution.— Northern Senegal and 
Mauritania across Western Sahara, Morocco, 
northern Algeria, southern Tunisia and north¬ 
western Libya, although absent from Mediter¬ 
ranean Morocco and Algeria (Sindaco and 
Jeremcenko 2008; Tamar et al. 2016b). The 
Libyan records mark the easternmost extent of 
the species. It was not included in the Libyan 
fauna by Schleich et al. (1996). 
Libyan Records (Map 27): TRIPOLITA- 
NIA: Nuqat al Khams : 7: Brito et al. 2008. 
Tripoli : 45: BMNH 1946.9.3.76; RMNH 
RENA 3493; Gray 1838, 1845; Boulenger 
1887, 1918, 1921; Crochet et al. 2003; Sindaco 
and Jeremcenko 2008. Nalut : 102: Brito et al. 
2008. 115: Brito et al. 2008. FEZZAN: Wadi 
al Shatii : 202: Brito et al. 2008. 203: Brito et 
al. 2008. Wadi al Hayaa : 268: Brito et al. 
2008. 
Map 27. Distribution of Acanthodactylus dumerilii in 
Libya. 
