BAUER, DEBOER, & TAYLOR: ATLAS OF THE REPTILES OF LIBYA 
213 
Comments. — Two subspecies are currently recognized within A. dumerilii by some authors 
(e.g., Salvador, 1982); the nominate form is West African, whereas the specimens found in North 
Africa, including Libya, belong to A. d. exiguus Lataste, 1885. Acanthodactylus dumerilii was syn- 
onymized with A. scutellatus by Boulenger (1887) and Arnold (1983), who considered the nature 
of variation in A. scutellatus to be too complex to warrant meaningful application of subspecific 
names. Sindaco and Jeremcenko (2008) did not recognize any subspecies in A. dumerilii. Tamar et 
al. (2016b) generated a multilocus phylogeny for the genus Acanthodactylus and found that A. 
dumerilii is made paraphyletic by A. senegalensis within the “ scutellatus clade.” Their sampling 
included no Libyan samples, however. Supposed A dumerilii from Senegal, Mauritania and West¬ 
ern Sahara in their study grouped with A. senegalensis, suggesting that these southwestern records 
for A. dumerilii need to be verified. The type locality for A. dumerilii is “Senegal” but, as noted by 
Crochet et al. (2003), this probably refers to an area in northwestern Africa outside of the nation of 
Senegal as currently recognized. Bons and Girot (1964) treated A. inornatus and 
A. dumerilii as different species and regarded the former as occurring across North Africa, although 
they plotted only two records in Libya, one in northwestern Tripolitania and the other along the 
Egyptian border and certainly referable to some congener. Crochet et al. (2003) showed that 
A. inornatus and A. exiguus are junior synonyms of A. dumerilii. 
IUCN Threat Status.— Not assessed, but assumed to be Least Concern. 
Acanthodactylus longipes Boulenger, 1918:154 
1918 Acanthodactylus scutellatus var. longipes Boulenger, Sur les lezards du genre Acanthodactylus 
Wieg. Bulletin de la Societe Zoologique de France 43:143-155. 
Lectotype.— BMNH 1946.8.4.31, “Wargla, Algeria.” designated by Crochet et al. (2003). Original pub¬ 
lished type locality “Sahara algerien.” 
Acanthodactylus scutellatus [part], Le Berre 1989:190. 
Acanthodactylus longipes , Schleich, Kastle, and Kabisch 1996:402. 
Acanthodactylus longipes, Sindaco and Jeremcenko 2008:220. 
Acanthodactylus longipes, Trape, Trape, and Chirio 2012:308. 
Distribution.— Mauritania, Mali and 
southern Morocco through Algeria, Niger, 
Libya and Chad to Egypt west of the Nile Val¬ 
ley and Delta (Tamar et al. 2016b). Libyan 
records are few and restricted to the inland west 
of Tripolitania and Fezzan, although the global 
distribution suggests that A. longipes should 
also inhabit the inland regions of eastern Libya. 
Libyan Records (Map 28): TRIPOLITA¬ 
NIA: Nalut : 125: Brito et al. 2008. FEZZAN: 
Wadi al Shatii : 209: Brito et al. 2008. Wadi al 
Hayaa : 277: Brito et al. 2008. Sabha : 285: 
BMNH 1954.1.6.42; Salvador 1982. Murzuo : 
298: BMNH (1) not cataloged; Salvador 1982. 
299: BMNH 1982.391. 312: Brito et al. 2008. 
“SW Libya”: photo, Sindaco and Jeremcenko 
2008; Wemer and Ibrahim 2012. 
Comments.— Baha El Din (1996) first 
Map 28. Distribution of Acanthodactylus longipes in 
Libya. 
