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Series 4, Volume 64, No. 8 
reported the species from Egypt, where it had previously been confused with A. scutellatus. Initial 
records were from across northern Egypt both east and west of the Nile and across northern Sinai 
(Baha El Din 1996). Specimens from east of the Nile Delta reported by Werner (1998) and Baha 
El Din (2006a) have since been allocated to A. aegyptius Baha El Din, 2007. Tamar et al. (2016b) 
generated a multilocus phylogeny for the genus Acanthodactylus and found A. longipes to be 
monophyletic and a member of the “scutellatus clade,” although they included no Libyan samples. 
As noted by Crochet et al. (2003), there are a small number of vouchered specimens given that the 
species has been mentioned by several authors as extremely common in Libya (Salvador, 1982; 
Schleich et. al. 1996; Sindaco and Jeremcenko 2008). Bons and Girot (1964) considered the species 
to be distributed across inland Tripolitania. 
IUCN Threat Status.— Not assessed, but assumed to be Least Concern. 
Acanthodactylus maculatus (Gray, 1838:281) (Fig. 22) 
Scapteira maculata Gray, Catalogue of the slender-tongued saurians, with descriptions of many new gen¬ 
era and species. Part 1. An nals and Magazine of Natural History, series 1, 1:274-283. 
Syntypes. — BMNH 1946.9.3.53 and two other specimens (fide Boulenger 1887, although Boulenger 
1921 mentioned only a single “type” and Salvador 19892 referred to the “Holotype”), “Tripoli” [Libya]. 
1918 Acanthodactylus pardalis Var. Latastei Boulenger, Sur les lezards du genre Acanthodactylus 
Wiegm. Bulletin de la Societe Zoologique de France 43:143-155. [153] 
Syntypes. — presumably BMNH, not located, “Tripoli, S. de la Tunisie, Algerie (Sahara et Hauts 
plateau).” 
Acanthodactylus scutellatus [part], Le Berre 1989:190. 
Acanthodactylus maculatus, Schleich, Kastle, and Kabisch 1996:392. 
Acanthodactylus maculatus, Sindaco and Jeremcenko 2008:218. 
Acanthodactylus maculatus, Trape, Trape, and Chirio 2012:310. 
Distribution.— From central Morocco and northern Algeria to Tunisia and Mediterranean 
Tripolitania (Tamar et al. 2016b). In Libya restricted to northwestern Tripolitania. 
Libyan Records (Map 29): TRIPOLITANIA: Zawiyah : 11: BMNH 1965.1256; Salvador 
1982. 12: MZUT R3265; ZMB 79710. 17: BMNH 1954.1.6.61,1955.1.8.66; Salvador 1982. 
Tripoli : 45: BMNH 1946.9.3.53; MZUT R2280; ZMB 14032, 15305, 15308-09, 15320-21, 
15383,18155; ZSM 484/1904; Gray 1838, 1845; Boulenger 1887,1918,1921;Konig 1888; Wern- 
Figure 22. Acanthodactylus maculatus from 5 km east of Al Jawsh, Nalut, Tripolitania, Libya, 2008. Photo © Roberto 
Sindaco. 
