BAUER, DEBOER, & TAYLOR: ATLAS OF THE REPTILES OF LIBYA 
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er 1909; Salvador 1982. Muroub : 68: BMNH 
1265.1258; Boulenger 1914; Salvador 1982. 
Misratah : 80: BMNH 1913.12.30.11; 
Boulenger 1914, 1921; Salvador 1982. Nalut : 
107: MCCI R1456; Sindaco, pers. obs. 
4/30/2008; Tamar et al. 2016b. 109: BMNH 
1965.1257-58; Salvador 1982. 118: BMNH 
1954.1.6.4; Salvador 1982. Jabal al Gharbi : 
136: BMNH 1965.1255; Salvador 1982. 154: 
ZSM 173/1979. 163: Brito et al. 2008; Fonseca 
et al. 2008. 164: Brito et al. 2008; Fonseca et al. 
2008. “Cyrenaica”: BMNH 1920.1.20.649 [in 
error, not plotted]. 
Comments.— Tamar et al. (2016b) gener¬ 
ated a multilocus phylogeny for the genus 
Acanthodactylus and identified A maculatus as 
a member of their “Western clade.” It is made 
paraphyletic by A. beershebensis, A. pardalis 
and A. bedriagai, although basal support values 
within their entire “ pardalis group” are too low to infer meaningful support for this pattern. Their 
sole Libyan sample was weakly supported as sister to conspecifics from Morocco and Algeria. The 
subspecific epithet “ latastei ” was subsequently misspelled “ latastii ” by Boulenger (1921). 
IUCN Threat Status.— Least Concern. 
Acanthodactylus pardalis (Lichtenstein, 1823: 99) 
1823 L[acerta ] Pardalis Lichtenstein, Verzeichniss der Doubletten des zoologischen Museums der 
Konigl. Universitat zu Berlin nebst Beschreibung vieler bisher unbekannter Arten von Saugethieren, Vogeln, 
Amphibien und Fischen. T. Trautwein, Berlin, x + 118 pp., 1 pi. 
Lectotype.— ZMB 1077, “Aegyptus”designated by Moravec et al. (1999). The paralectotypes are ZMB 
1072-1076 and ZMB 55905 (formerly part of ZMB 1077) (Bauer and Gunther 1995). 
Acanthodactylus pardalis, Le Berre 1989:188. 
Acanthodactylus pardalis, Schleich, Kastle, and Kabisch 1996:388. 
Acanthodactylus pardalis, Sindaco and Jeremcenko 2008:218. 
Distribution.— Mediterranean coast from the Gulf of Sirte east to the Nile Valley of Egypt 
(Sindaco and Jeremcenko 2008; Tamar et al. 2016b), a single record from east of the Nile is dis¬ 
cussed by Moravec et al. (1999). In Libya they are found along the coast from the eastern Gulf of 
Sirte to the Egyptian border, but are absent from the elevated regions of the Jabal al Akhdar. 
Libyan Records (Map 30): CYRENAICA: Benghazi : 342: MZUT R2289; Calabresi 1923; 
Zavattari 1929, 1930, 1934. 355: BMNH 1954.1.6.5-7, Salvador 1982. 357: BMNH 1988.189, 
1988.191-92; MZUF 852, 12688; NHMW 8161; von Martens 1883; Werner 1909; Ghigi 1920; 
Calabresi 1923; Umani 1923; Zavattari 1929, 1930, 1934; Moravec et al. 1999. 358: MZUF 853; 
Calabresi 1923; Zavattari 1929, 1930, 1934; Moravec et al. 1999. 364: Comalia in Haimann 1882, 
1886; Zavattari 1934. 367: MZUF 770; MZUT R2412; Calabresi 1923; Zavattari 1929, 1930, 
1934; Moravec et al. 1999. 376: BMNH 1965.1259; Salvador 1982. Marj : 385: MCSN 2079; 
Zavattari 1937. 399: Fonseca et al. 2008. 402: MZUT R2297; Calabresi 1923; Zavattari 1929, 
1930. 403: NHMC 80.3.73.1. Jabal al Akhdar : 417: Zavattari 1929, 1930. 452: Ghigi 1920; Cal- 
Map 29. Distribution of Acanthodactylus maculatus in 
Libya. 
