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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 8 
32958-69; MZUT R1942; Calabresi 1923; Zavattari 1929, 1930. 376: BMNH 1965.1191. Marj : 
385: BMNH 1965.1190; MCSN 2192-93, 2204, 2322; SMF 58325; Ghigi 1920; Calabresi 1923; 
Zavattari 1929, 1930, 1934, 1937; Schleich 1987; Greenbaum et al. 2006. 388: NHMC 
80.3.80.976-977; Komilios et al. 2010. 389: MZUT R1937; Zavattari 1922, 1929, 1930, 1934; 
Calabresi 1923; Frynta et al. 2000. 398: NHMC 80.3.80.41, 80.3.80.43; Komilios et al. 2010. 403: 
NHMC 80.3.80.37-38; Komilios et al. 2010. 408: MZUT R1917; Calabresi 1923; Zavattari 1929, 
1930. 457ai: Schleich 1987. 457b: Schleich 1987. 457v: Schleich 1987. Jabal al Akhdar : 416: 
NHMC 80.3.80.33-35; Komilios et al. 2010. 417: MCSN 2468; SMF 34463; Zavattari 1922,1929, 
1930, 1934; Calabresi 1923; Frynta et al. 2000. 419: FMNH 214963; Resetar 1981. 420: NHMC 
80.3.80.32, 80.3.80.722-723. 421: Vinciguerra 1927; Zavattari 1929, 1930, 1934; Gestro and Vin- 
ciguerra 1931. 428: ZSM 125/1983[5 specimens]; Schleich 1987. 432: FMNH 214961; Resetar 
1981. 434: FMNH 83060, 214962; Resetar 1981. 436: NHMC 80.3.80.36; Komilios et al. 2010. 
440: FMNH 214960; Resetar 1981. 452: Ghigi 1920; Calabresi 1923; Zavattari 1929, 1930, 1934. 
457: FMNH 83059; KNP 1981/172-173, 194-195, 242, 255, 257, 286, 362, 365, 405, 411, 424- 
427, 433^144, 454, 481, 491, 495, 499; SMNS 1094; ZSM 458-62/1997; Schleich 1987. 457at: 
Schleich 1987. 457au: Schleich 1987. 457av: Schleich 1987. 457aw: Schleich 1987. 457ay: Schle¬ 
ich 1987. 457ba: Schleich 1987. 457bb: Schleich 1987. 457bc: Schleich 1987. 457be: Schleich 
1987. 457bi: Schleich 1987. 457bj: Schleich 1987. 457bq: Schleich 1987. 457bt: Schleich 1987. 
457ch: Schleich 1987. 457cs: Schleich 1987. 457cu: Schleich 1987. 4571: Schleich 1987. Darnah : 
458: NHMC 80.3.80.40. 460: NHMC 80.3.80.39; Komilios et al. 2010. 462: Zavattari 1929, 1930. 
463: Calabresi 1923; Zavattari 1929, 1930, 1934. 465: MZUT R1913; Calabresi 1923; Zavattari 
1929,1930, 1934. 243c: Calabresi 1923. 466: MZUT R1920, R1933; SMF 34473; NHMW 10444; 
Werner 1909; Andreucci 1913; Ghigi 1913, 1920; Calabresi 1923; Zavattari 1929, 1930, 1934; 
Greenbaum et al. 2006. 467: Zavattari 1929, 1930. 472: NHMC 80.3.80.42; Komilios et al. 2010. 
475: SMNS 1083. 477: NHMC 80.3.80.44^17; Komilios et al. 2010. Butnan : 490: NHMC 
80.3.80.48; Komilios et al. 2010; SMF 54085; ZMB 62938; Zavattari 1922,1929,1930,1934; Cal¬ 
abresi 1923; Vinciguerra 1927; Gestro and Vinciguerra 1931. 493: NHMC 80.3.80.49. 494: ZSM 
124/1983. 504: Vinciguerra 1927; Zavattari 1929, 1930; Gestro and Vinciguerra 1931. 505: Vin¬ 
ciguerra 1927; Zavattari 1929, 1930, 1934; Gestro and Vinciguerra 1931. 517: Zavattari 1929, 
1930. Al Wahat : 544: BMNH 1965.1189; SMF 58326; Vinciguerra 1931; Zavattari 1934; Green¬ 
baum et al. 2006. 553: MCSN 2184, Zavattari 1934, 1937. 562: ZMB 17999; Peters 1880, 1881; 
Werner 1909; Andreucci 1913; Ghigi 1913; Zavattari 1929, 1930, 1934. 564: BMNH 1932.3.6.15; 
MCSN 2195; Zavattari 1929, 1930, 1937; Vinciguerra 1931. Kufrah : 568: Zavattari 1937. 571: 
MZUF 861; Scortecci 1935c. 573: Vinciguerra 1931; Zavattari 1934, 1937. 575: BMNH 
1932.3.6.13-14; Vinciguerra 1931. 575a: Vinciguerra 1931. “Cyrenaica”: BMNH 1954.1.7.20- 
24; ZMB 10500, 74194. “Marmarica”: Zavattari 1937. “Nordost-Libyen”: ZMB 62931-37, 
62948-54. LIBYA: ZMB 18002. “deserto di Sahara”: Rizzardi 1896. 
Comments.— Greenbaum et al. (2006) revalidated many sub-Saharan African congeners, in 
addition to synonymizing the central Saharan C. ragazzii Boulenger 1890 with C. o. ocellatus (Sin- 
daco and Jeremcenko 2008). Schleich et al. (1996) and Sindaco and Jeremcenko (2008) recognized 
five subspecies, with only the nominate form in Libya. The nominate form occupies nearly all of 
the species’ range except portions of northeast Morocco and northwest Algeria (C. o. subtypicus 
Werner, 1931), Mediterranean areas of North Africa from northeast Morocco to Tunisia as well as 
Sicily and Sardinia (C. o. tiligugu (Gmelin, 1789)), and Lampedusa and Lampione Islands (C. o. 
zavattarii Lanza, 1954) and Linosa Island (C. o. linosae Boulenger, 1920). Komilos et al. (2010) 
found members of three different lineages in Libya, one in northwestern Tripolitania shared with 
most of Tunisia, Sardinia, Sicily and Malta, one occupying the north of Tripolitania to the western 
