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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 8 
MSNG 29517. 220: MCSN 2790; Kramer and Schnurrenberger 1963. 229: SK 1897-98; Kramer 
and Schnurrenberger 1963. Sabha : 289: SK 1445; Kramer and Schnurrenberger 1963. Murzuq : 
296: ZCT 2005.30; Ibrahim 2008a. CYRENAICA: Benghazi : 367: CAS 135739. 377: SK 48; 
Kramer and Schnurrenberger 1963. 378: SK 381, 571, 752. 379: Kramer and Schnurrenberger 
1958. Marj : 385: Werner 1909. Jabal al Akhdar : 423: CM 91594. Darnah : 475: ANSP 28051. 
477: NHMC 80.3.23.2. Butnan : 508: NHMC 80.3.23.1. 512: ZSM 158/1983. AlWahat : 536: SK 
329; Kramer and Schnurrenberger 1963. 544: SMF 62312. 547: SK 66-67; Kramer and Schnur¬ 
renberger 1958,1963. 548: AMNH R109610; SK 977-78; Kramer and Schnurrenberger 1963. 549: 
SK 401-04; Kramer and Schnurrenberger 1963. 562: MSNG 31586; Vinciguerra 1931; Zavattari 
1934; Scortecci 1939. Kufrah : 575: Peters 1880, 1881; Wemer 1909; Ghigi 1913, Zavattari 1929, 
1930, 1934; Scortecci 1939. LIBYA: NHMW-25664. 
Comments.— Bohme and de Pury (2011) discussed the allocation of this species to different 
genera and presented evidence for its inclusion in Rhagerhis. Most recently, however, Figueroa et 
al. (2016) presented evidence that R. moilensis was nested within Malpolon and transferred the 
species back to this genus. Pending further study we here retain it in Rhagerhis. 
UCN Threat Status: Not assessed, but assumed to be Least Concern. 
Family Elapidae 
Naja haje (Linnaeus, 1758:225) 
1758 Coluber Haje Linnaeus, Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Gen¬ 
era, Species, cum Characteribus, Differentiis, Synonymis, Locis. Tomus I. Editio decima, reformata. Lauren- 
tii Salvii, Holmiae [Stockholm], Sweden. (4) + 823 + (I) pp. 
Syntypes. — NRM and Hasselquist collection (see Comments), lost fide Andersson (1899) and Kramer 
and Schnurrenberger (1963), “/Eegypto inferiore” [= Lower Egypt], 
Naja haje, Kramer and Schnurrenberger 1963:536. 
Naja haje, Le Berre 1989:256. 
Naja haje, Schleich, Kastle, and Kabisch 1996:526. 
Naja haje, Sindaco, Venchi and Grieco 2013:176. 
Distribution.— Across northern Africa exclusive of the Sahara proper, from Senegal to 
Western Sahara and Morocco to the Nile Valley and Delta (Baha El Din 2006a). In the Sahel cross¬ 
ing eastwards through northern Nigeria, southern Chad and the Central African Republic into 
Sudan and the Horn of Africa and as far south as Tanzania in East Africa. Libyan records are chiefly 
in coastal or near-coastal regions across the country. 
Libyan Records (Map 62): TRIPOLITANIA: Tripoli : 45: Wemer 1909; Scortecci 1939. 
Muroub : 60: Wemer 1909; Ghigi 1913; Zavattari 1934; Scortecci 1939. 68: SK 448; Kramer and 
Schnurrenberger 1963. Jabal al Gharbi : 147: FMNH 83058. “Tripolitania settentrionale” 
Zavattari 1937. Sirte : 174: MCSN 2688. CYRENAICA: Benghazi : 354: Umani 1922. 357: von 
Martens 1883; Wemer 1909; Umani 1922; Zavattari 1922, 1929, 1930, 1934; Calabresi 1923; 
Ribolla 1923; Scortecci 1939. 376: Zavattari 1922, 1929, 1930, 1934; Calabresi 1923; Scortecci 
1939. Marj : 385: Zavattari 1929, 1937. Jabal al Akhdar : 411: FMNH 214914; Resetar 1981. 
414: Resetar 1981. 417: Zavattari 1922, 1929, 1930, 1934; Calabresi 1923; Scortecci 1939. 421: 
Ghigi 1920; Calabresi 1923; Zavattari 1929, 1930. 427: Scortecci 1939. 457: KNP 1981/272, 354, 
509; ZSM 145/1983; Resetar 1981; Schleich 1987. 457aj: Schleich 1987. 457an: Schleich 1987. 
457ao: Schleich 1987. 457at: Schleich 1987. 457au: Schleich 1987. 457bp: Schleich 1987. 457br: 
Schleich 1987. 457bs: 179: KNP 1919/13; Schleich 1987. 457bu: Schleich 1987. 457ca: Schleich 
