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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 8 
Jabalal Gharbi : 131: SK 1750,2458; Kramer 
and Schnurrenberger 1963._158: MCSN 2927. 
Sirte : 185: MCSN 2929; Kramer and Schnur¬ 
renberger 1963. “Tripolitania”: Scortecci 
1939. “Tripolitania settentrionale” Zavattari 
1937. FEZZAN: Ghat : 233: Schnurrenberger 
1959. 239: Scortecci 1937a. 245: Scortecci 
1937a. 249: Scortecci 1939. Murzuo : 298: 
MCSN 2946, 2958. 305: ZCT 2005.36; 
Ibrahim 2008a. 316: ZCT 2006.06; Ibrahim 
2008a. “Ghat-Murzuq-Mesak-Mellet-Uadi 
Tilemsin”: MCSN 2933. CYRENAICA: 
Benghazi : 358: MZUF 989. 372: Frynta et al. 
2000; Kratochvil et al. 2002. Mare 384: CM 
91617. Butnan : 491: MZUF 992. 521: MSNG 
37503; Zavattari 1929, 1930, 1934, 1937. Al 
Wahat : 536: SK 323, 423-24, 287; Kramer 
and Schnurrenberger 1963. 540: Schnurren¬ 
berger 1959. 544: BMNH 1960.1.6.11-12; CM S7209; Kramer and Schnurrenberger 1963; BMNH 
1985.1181. 547: Kramer and Schnurrenberger 1958. 548: SK 301-05; Kramer and Schnurrenberg¬ 
er 1963. 549: SK 405-08, 630; Kramer and Schnurrenberger 1963. 550: Schnurrenberger 1958a. 
553: MCSN 4198, Zavattari 1937. 554: MCSN 2965; Scortecci 1935b; Kramer and Schnurren¬ 
berger 1963. 562: MSNG 31581*; Vinciguerra 1931; Zavattari 1934. 564: MSNG 31581*; Vin- 
ciguerra 1931; Zavattari 1934, 1937. Kufrah : 575: SMF 32710; ZMB 18091; Scortecci 1939. 
“Cyrenaica”: MCSN 2959; MZUF 12153-54; Gestro and Vinciguerra 1931; Scortecci 1939. 
“Soluch to Hatiet el Rtem”: Schnurrenberger 1959. 
Comments.— Wagner and Wilms (2010) commented on the conflicting reports of sympatry 
and syntopy in the literature regarding the spatial relationships between C. cerastes and C. vipera. 
Our records support that the two occur in sympatry, although C. vipera tends to occupy more xeric 
environments than C. cerastes, as was concluded by Wagner and Wilms (2010). Schnurrenberger 
(1963) and Kramer and Schnurrenberger (1963) noted that the localities provided for the two 
species of Cerastes by Schnurrenberger (1959) were reversed. Scortecci (1937a) listed several 
localities from near Ghat including Tafilellelt, Uadi Iseien near Tin Caraden, Uadi Inelegghi, and 
Uadi Iseien near Tin Alcun. Kramer and Schnuerrenberger (1963) treated the first two of these as 
being in Algerian territory, but considered the other two to be Libyan. However, all four of these 
localities are actually in the Tassili n’Ajjer of Algeria. Likewise, Scortecci’s (1939) mention of a 
specimen from the Tassili certainly refers to an Algerian snake. 
IUCN Threat Status: Least Concern. 
Echis pyramidum (I. Geoffroy Saint-Hilaire, 1827:151, pi. 8, fig. 1) (Fig. 36) 
1827 Scythalepyramidum Geoffroy Saint-Hilaire, Description des reptiles qui se trouvent en Egypte, pp. 
121-160 in M.J.-C.L. de Savigny, (ed.), Description de l’Egypte, ou Recueil des Observations et des 
Recherches qui ont ete Faites en Egypte Pendant l Expedition de l ’Armee Frangaise. Histoire Naturelle. Tome 
premier. Partie premier. Imprimerie Imperiale, Paris, France. 
Lectotype. — MNHN 4031, designated by Stemmier and Sochurek (1969:90), “Kaire, Egypte” [= Cairo, 
Egypt], 
1972 Echis carinatus leucogaster Roman, Deux sous-especes de la vipere Echis carinatus (Schneider) 
Map 64. Distribution of Cerestes vipera in Libya. 
