BAUER, DEBOER, & TAYLOR: ATLAS OF THE REPTILES OF LIBYA 
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dans les territories de Haute-Volta et du Niger: Echis carinatus ocellatus Stemmier — Echis carinatus leuco¬ 
gaster n. ssp. Notes et Documents Voltaiques, 5(4): 1-11, 2 maps, 1 pi. [6, pi. figs. 3-8]. 
Holotype.— RN 191, “Boubon, a 20 km au Nord de Niamey” [Niger]. 
1990 Echis pyramidum lucidus Cherlin, Taxonomic revision of the snake genus Echis (Viperidae). II. An 
analysis of taxonomy and description of new forms [in Russian]. Trudy Zoologicheskogo Instituta Akademii 
NaukSSSR, 207:193-223, pis. IV-VI. [206, pi. V, fig. 3], 
Holotype.— USNM 134975, “ErnneT, OaioM, CmniypHc” [Egypt, Faiyum, Sinnuris]. 
Echis carinatus pyramidum, Kramer and Schnurrenberger 1963:544. 
Echis leucogaster, Le Berre 1989:298. 
Echis pyramidum, Le Berre 1989:300. 
Echis pyramidum lucidus, Schleich, Kastle, and Kabisch 1996:542. 
Echis arenicola leucogaster, Schleich, Kastle, and Kabisch 1996:540. 
Echis pyramidum leucogaster, Sindaco, Venchi, and Grieco 2013:200. 
Figure 36. Echis pyramidum from between A1 Khums and Tripoli, Tripolitania, Libya (32.6694, 14.1242), 2007. Photo 
courtesy of Natural History Museum of Crete © V. Paravas. 
Distribution.— From Senegal and Guinea-Bissau across North Africa to Egypt and south to 
Kenya and the Horn of Africa. In both Egypt and Libya there are scattered localities, none from 
Kufrah and the southeastern portion of the country, nor from the Egyptian border region. 
Libyan Records (Map 65): TRIPOLITANIA: Tripoli : 45: BMNH 1960.1.6.13; Kramer and 
Schnurrenberger 1963. Muroub : 65: NHMC 80.3.4.1. 68: Boulenger 1914; Zavattari 1934; 
Scortecci 1939. “Tripolitania settentrionale”: Zavattari 1937. FEZZAN: Sabha : 286: NMBA- 
REPT 18127. CYRENAICA: Butnan : 490: Ribolla 1923; Zavattari 1934; Scortecci 1939. “Cyre- 
naica”: Condorelli-Francaviglia 1896; Schleich et al. 1996; Sindaco et al. 2013; Werner 1909; Cal- 
abresi 1923. “Marmarica”: Zavattari 1937. 
Comments.— The complicated dating of the natural history portions of Description de 
