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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, No. 8 
Table 2. Distribution of the reptile species and subspecies recorded in Libya. The first three columns 
show occurrence in the three regions of Libya: Tripolitania (TRP), Fezzan (FEZ), and Cyrenaica (CYR); the 
remaining columns shows their distribution in the six countries sharing borders with Libya: Egypt (EGY), 
Sudan (SUD), Chad (CHA), Niger (NIG), Algeria (ALG), and Tunisia (TUN). Data for Libya from the pres¬ 
ent study, that for other North African countries from Sindaco and Jeremcenko (2008), Trape et al. (2012), and 
Sindaco et al. (2013) with updates from the relevant taxonomic literature. I = introduced, M = marine species, 
N/A = not applicable, X = species occurrence confirmed, ?: species questionably or possibly present (see cor¬ 
responding species account). Note that Chad, in particular, remains very poorly sampled, thus absence of con¬ 
firmation does not necessarily imply a lack of actual occurrence. 
LIBYA_Neighboring Countries 
Taxon 
TRP 
FEZ 
CYR 
EGY 
SUD 
CHA 
NIG 
ALG 
TUN 
CHELONIA 
Chelonudae 
Caretta caretta 
M 
N/A 
M 
M 
N/A 
N/A 
M 
M 
Chelonia mydas 
N/A 
M 
M 
M 
N/A 
N/A 
M 
M 
Dermochelyidae 
Dermochelys coriacea 
M 
N/A 
M 
M 
N/A 
N/A 
M 
M 
Geomydidae 
Mauremys leprosa saharica 
X 
X 
X 
Testudinidae 
Testudo graeca nabeulensis 
X 
X 
Testudo graeca cyrenaica ENDEMIC 
X 
Testudo kleinmanni 
X 
X 
X 
SQUAMATA 
Agamidae 
Agama tassiliensis 
X 
X 
X 
X 
Pseudotrapelus chlodnickii 
X 
X 
X 
7 
Trapelus mutabilis mutabilis 
X 
X 
X 
X 
X 
X 
X 
Trapelus mutabilis poppeki ENDEMIC 
X 
X 
Uromastycidae 
Uromastyx acanthinura 
X 
X 
X 
X 
X 
Uromastyx alfredschmidti 
X 
X 
Chamaeleonidae 
Chamaeleo chamaeleon 
X 
X 
X 
X 
X 
X 
Gekkonidae 
Hemidactylus turcicus 
X 
X 
X 
X 
X 
X 
Stenodactylus mauritanicus 
X 
X 
X 
X 
X 
Stenodactylus petrii 
X 
X 
X 
X 
X 
X 
X 
X 
Stenodactylus stenurus 
X 
X 
X 
Stenodactylus sthenodactylus 
X 
X 
X 
X 
X 
X 
X 
X 
X 
Tropiocolotes steudneri 
X 
X 
X 
X 
X 
Tropiocolotes tripolitanus 
X 
X 
X 
X 
X 
X 
Phyllodactylidae 
Ptyodactylus togoensis 
X 
X 
X 
X 
Tarentolci annularis 
X 
X 
X 
X 
X 
X 
Tarentola deserti 
X 
X 
X 
Tarentola fascicularis Complex 
X 
X 
X 
X 
X 
X 
Tarentola mindiae 
X 
X 
Tarentola neglecta neglecta 
X 
X 
X 
X 
Tarentola neglecta lanzai ENDEMIC 
X 
Lacertidae 
Acanthodactylus boskianus asper 
X 
X 
X 
X 
X 
X 
X 
X 
X 
Acanthodactylus dumerilii 
X 
X 
X 
X 
Acanthodactylus longipes 
X 
X 
7 
X 
X 
X 
X 
X 
Acanthodactylus maculatus 
X 
X 
X 
Acanthodactylus pardalis 
X 
X 
Acanthodactylus scutellatus audouini 
X 
X 
X 
X 
X 
X 
X 
X 
X 
Mesalina guttulata 
X 
X 
X 
X 
X 
X 
X 
X 
