ELLIOTT ET AL.: THE CAVE FAUNA OF CALIFORNIA 
67 
Overall, 105 cave obligates occupy 291 caves (36% of sampled caves). Occupancy provides 
some measure of the success in finding obligate species in each region, and the potential for pro¬ 
ductive future work. Occupancy is high in the Sierra Nevada South (SNS) (0.64) and moderate in 
the Sierra Nevada North (SNN) (0.41). This also reflects the high endemism and biodiversity with¬ 
in SNS caves, discussed below. 
Table 11. List of 146 obligate subterranean species in California. TB is troglobite, SB is stygobite, and 
PB is phreatobite. Twelve species are designated TB?, indicating that the status as a troglobite is not yet 
certain. 
Species 
Adaptation 
Class 
Order 
Family 
Total Sites 
Amplaria shastae 
TB 
Diplopoda 
Chordeumatida 
Striariidae 
2 
Anillaspis explanata 
TB 
Insecta 
Coleoptera 
Carabidae 
1 
Anillina n. gen., n. sp. no. 1 
TB 
Insecta 
Coleoptera 
Carabidae 
2 
Anillina n. gen., n. sp. no. 2 
TB 
Insecta 
Coleoptera 
Carabidae 
1 
Anillina n. gen., n. sp. no. 3 
TB 
Insecta 
Coleoptera 
Carabidae 
1 
Anillina n. gen., n. sp. no. 4 
TB 
Insecta 
Coleoptera 
Carabidae 
2 
Aphrastochthonius grubbsi 
TB 
Arachnida 
Pseudoscorpiones 
Chthoniidae 
1 
Aphrastochthonius similis 
TB 
Arachnida 
Pseudoscorpiones 
Chthoniidae 
1 
Apochthonius grubbsi 
TB 
Arachnida 
Pseudoscorpiones 
Chthoniidae 
4 
Australinocreagris grahami 
TB 
Arachnida 
Pseudoscorpiones 
Neobisiidae 
15 
Australinocreagris sp. 
TB 
Arachnida 
Pseudoscorpiones 
Neobisiidae 
4 
Banksula californica 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Banksula galilei 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Banksula grahami 
TB 
Arachnida 
Opiliones 
Phalangodidae 
18 
Banksula grubbsi 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Banksula martinorum 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Banksula rudolphi 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Banksula tuolumne 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Banksula tutankhamen 
TB 
Arachnida 
Opiliones 
Phalangodidae 
1 
Bathynella fraterna 
PB 
Malacostraca 
Bathynellacea 
Bathynellidae 
1 
Bathynella germanitas 
PB 
Malacostraca 
Bathynellacea 
Bathynellidae 
1 
Bidentogon n. sp. 
TB 
Diplopoda 
Polydesmida 
Polydesmidae 
5 
Blabomma n. sp. 
TB 
Arachnida 
Araneae 
Dictynidae 
1 
Blabomma n. sp. no. 1 
TB 
Arachnida 
Araneae 
Dictynidae 
4 
Blabomma n. sp. no. 2 
TB 
Arachnida 
Araneae 
Dictynidae 
6 
Blabomma n. sp. no. 3 
TB 
Arachnida 
Araneae 
Dictynidae 
1 
Blabomma n. sp. no. 4 
TB 
Arachnida 
Araneae 
Dictynidae 
1 
Blabomma sp. 
TB 
Arachnida 
Araneae 
Dictynidae 
3 
Bowmanasellus sequoiae 
SB 
Malacostraca 
Isopoda 
Asellidae 
9 
Caecidotea n. sp. 
SB 
Malacostraca 
Isopoda 
Asellidae 
1 
Calasellus californicus 
PB 
Malacostraca 
Isopoda 
Asellidae 
24 
Calasellus longus 
PB 
Malacostraca 
Isopoda 
Asellidae 
1 
Calasellus sp. 
PB 
Malacostraca 
Isopoda 
Asellidae 
13 
Calicina cloughensis 
TB 
Arachnida 
Opiliones 
Phalangodidae 
4 
Califobathynella noodti 
PB 
Malacostraca 
Bathynellacea 
Parabathynellidae 
1 
