ELLIOTT ET AL.: THE CAVE FAUNA OF CALIFORNIA 
69 
Table 11 (continued). List of 146 obligate subterranean species in California. TB is troglobite, SB is 
stygobite, and PB is phreatobite. Twelve species are designated TB?, indicating that the status as a troglobite 
is not yet certain. 
Species 
Adaptation 
Class 
Order 
Family 
Total 
Sites 
Neochthonius n. sp. 
TB 
Arachnida 
Pseudoscorpiones 
Chthoniidae 
3 
Neochthonius sp. 
TB 
Arachnida 
Pseudoscorpiones 
Chthoniidae 
4 
Neochthonius troglodytes 
TB 
Arachnida 
Pseudoscorpiones 
Chthoniidae 
10 
Nesticus potterius 
TB 
Arachnida 
Araneae 
Nesticidae 
4 
Nevadesmus n. sp. 1 
TB 
Diplopoda 
Polydesmida 
Macrostemodesmidae 
1 
Nevadesmus n. sp. 2 
TB 
Diplopoda 
Polydesmida 
Macrostemodesmidae 
1 
Niptus arcanus 
TB 
Insecta 
Coleoptera 
Anobiidae 
1 
Oaphantes n. sp. 1 
TB 
Arachnida 
Araneae 
Linyphiidae 
1 
Occasjapyx kofoidi 
TB 
Diplura 
Dicellurata 
Japygidae 
2 
Opiona graeningi 
TB 
Diplopoda 
Chordeumatida 
Caseyidae 
11 
Opiona n. sp. no. 2 
TB 
Diplopoda 
Chordeumatida 
Caseyidae 
1 
Pacificabathynella sequoiae 
PB 
Malacostraca 
Bathynellacea 
Bathynellidae 
1 
Parobisium yosemite 
TB 
Arachnida 
Pseudoscorpiones 
Neobisiidae 
2 
Plumatyla humerosa 
TB 
Diplopoda 
Chordeumatida 
Conotylidae 
43 
Plumatyla sp. 
TB 
Diplopoda 
Chordeumatida 
Conotylidae 
2 
Polycelis sp. 
SB? 
Turbellaria 
Tricladida 
Planariidae 
6 
Pratherodesmus despaini 
TB 
Diplopoda 
Polydesmida 
Macrostemodesmidae 
1 
Pratherodesmus n. sp. 
TB 
Diplopoda 
Polydesmida 
Macrostemodesmidae 
1 
Prokoenenia sp. 
TB 
Arachnida 
Palpigradi 
Prokoeneniidae 
2 
Pseudogarypus orpheus 
TB 
Arachnida 
Pseudoscorpiones 
Pseudogarypidae 
2 
Pseudogarypus spelaeus 
TB 
Arachnida 
Pseudoscorpiones 
Pseudogarypidae 
1 
Ptomaphagus ( Adelops ) inyoensis 
TB 
Insecta 
Coleoptera 
Leiodidae 
1 
Sequoiadesmus krejcae 
TB? 
Diplopoda 
Polydesmida 
Macrostemodesmidae 
3 
Sinella tecta 
TB 
Collembola 
Collembola 
Entomobryidae 
5 
Speleonycta n. sp. no. 1 
TB 
Insecta 
Zygentoma 
Nicoletiidae 
1 
Speleonycta n. sp. no. 2 
TB 
Insecta 
Zygentoma 
Nicoletiidae 
4 
Speoseya grahami 
TB 
Diplopoda 
Chordeumatida 
Caseyidae 
3 
Sphalloplana ( Sphalloplana ) californica 
SB 
Turbellaria 
Tricladida 
Kenkiidae 
1 
Spirembolus sp. 
TB 
Arachnida 
Araneae 
Linyphiidae 
2 
Stenophilus californicus 
TB? 
Chilopoda 
Geophilomorpha 
Himantariidae 
2 
Stygobromus cherylae 
PB 
Malacostraca 
Amphipoda 
Crangonyctidae 
1 
Stygobromus cowani 
PB 
Malacostraca 
Amphipoda 
Crangonyctidae 
1 
Stygobromus gallawayae 
PB 
Malacostraca 
Amphipoda 
Crangonyctidae 
1 
Stygobromus gradyi 
SB 
Malacostraca 
Amphipoda 
Crangonyctidae 
5 
Stygobromus grahami 
SB 
Malacostraca 
Amphipoda 
Crangonyctidae 
8 
Stygobromus harai 
SB 
Malacostraca 
Amphipoda 
Crangonyctidae 
4 
Stygobromus hyporheicus 
PB 
Malacostraca 
Amphipoda 
Crangonyctidae 
1 
