ELLIOTT ET AL.: THE CAVE FAUNA OF CALIFORNIA 
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Table 18. Clough Cave, Sequoia National Park, Tulare County, Sierra Nevada South Region, species list 
(58 species). Site endemism (SE) scores are for troglobites only, based on the number of sites statewide. 
Phylum/ Class 
Order 
Family 
Species 
Ecological 
classification 
SE 
Arthropoda 
Arachnida 
Acari 
Undetermined 
Acari 
Araneae 
Anapidae 
Gertschanapis shantzi 
Troglophile 
Cybaeidae 
Cybaeus sp. 
Epigean 
Dictynidae 
Blabomma n. sp. no. 4 
Troglobite 
1.00 
Dictynidae 
Yorima n. sp. 
Troglobite 
0.11 
Hahniidae 
Calymmaria sp. 
Epigean 
Hypochilidae 
Hypochilus petnmkevitchi 
Epigean 
Leptonetidae 
Archoleptoneta schusteri 
Troglophile 
Leptonetidae 
Calileptoneta n. sp. 2 
Troglobite 
0.25 
Linyphiidae 
Arcuphantes sp. 
Troglophile 
Linyphiidae 
Linyphiidae 
Nesticidae 
Nesticus silvestrii 
Troglophile 
Pholcidae 
Physocyclus sp. 
Troglophile 
Telemidae 
Usojila n. spp. 
Troglobite 
0.06 
Tengellidae 
Titiotus sp. 
Troglophile 
Undetermined 
Araneomorphae 
Opiliones 
Phalangodidae 
Calicina clonghensis 
Troglobite 
0.25 
Pseudoscorpiones 
Chthoniidae 
Neochthonius n. sp. 
Troglobite 
0.33 
Chthoniidae 
Neochthonius sp. 
Uncertain 
Neobisiidae 
Fissilicreagris sp. 
Troglobite 
0.50 
Schizomida 
?Hubbardiidae 
1 Hubbardia sp. 
Scorpiones 
Undetermined 
Scorpiones 
Vaejovidae 
Uroctonites sequoia 
Troglophile 
Vaejovidae 
Uroctonus mordax 
Epigean 
Chilopoda 
Lithobiomorpha 
Undetermined 
Lithobiomorpha 
Uncertain 
Undetermined 
Undetermined 
Chilopoda 
Collembola 
Collembola 
Sminthuridae 
Sminthuridae 
Tomoceridae 
Pogonognathellus celsus 
Troglophile 
Tomoceridae 
Tomoceridae 
Undetermined 
Collembola 
Diplopoda 
Chordeumatida 
Conotylidae 
Taiyutyla sp. 
Troglophile 
Striariidae 
Amplaria adamsi 
Troglophile 
Julida 
Paeromopodidae 
Californiulus yosemitensis 
Endogean 
Polydesmida 
Macrostemodesmidae 
Sequoiadesmus krejcae 
Troglobite? 
0.33 
Polydesmidae 
Bidentogon n. sp. 
Troglobite 
0.20 
Undetermined 
Undetermined 
Diplopoda 
Diplura 
Dicellurata 
Japygidae 
Japygidae 
Cave-limited 
Insecta 
Archaeognatha 
Undetermined 
Archaeognatha 
Coleoptera 
Carabidae 
Carabidae 
Monotomidae 
Hesperobaenus abbreviatus 
Raymondionymidae 
Gilbertiola sp. 
Troglobite 
1.00 
Undetermined 
Coleoptera 
Diptera 
Mycetophilidae 
Mycetophilidae 
Undetermined 
Diptera 
Hemiptera 
Achilidae 
Juniperthia succinea 
Aphididae 
Aphididae 
Psocoptera 
Psyllipsocidae 
Psyllipsocus ramburii 
Troglophile 
Undetermined 
Psocoptera 
Undetermined 
Undetermined 
Insecta 
Zygentoma 
Nicoletiidae 
Speleonycta n. sp. no. 1 
Troglobite 
1.00 
