84 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, Supplement 1 
Table 19. Samwel Cave, Shasta County, Shasta National Forest, Klamath Mountains Region, species list 
(44 species). 
Phylum/Class 
Order 
Family 
Species 
Ecological 
classification 
SE 
Arthropoda 
Arachnida 
Araneae 
Amaurobiidae 
Callobius tehama 
Epigean 
Amaurobiidae 
Pimus nawtawaketus 
Troglophile 
Cybaeidae 
Cybaeozyga n. sp. 2 
Troglobite 
1.00 
Cybaeidae 
Cybaeus n. sp. 
Troglobite 
1.00 
Leptonetidae 
Calileptoneta sylva 
Troglophile 
Nesticidae 
Nesticus potterius 
Troglobite 
0.25 
Telemidae 
Usofila n. spp. 
Troglobite 
0.06 
Tengellidae 
Titiotus humboldt 
Troglophile 
Pseudoscorpiones 
Chthoniidae 
Neochthonius sp. 
Uncertain 
Pseudogarypidae 
Pseudogarypus spelaeus 
Troglobite 
1.00 
Scorpiones 
Vaejovidae 
Graemeloweus iviei 
Troglophile 
Vaejovidae 
Uroctonus grahami 
Troglobite 
1.00 
Chilopoda 
Lithobiomorpha 
Undetermined 
Lithobiomorpha 
Uncertain 
Collembola 
Collembola 
Dicyrtomidae 
Ptenothrix maculosa 
Entomobryidae 
Entomobryoides guthriei 
Troglophile 
Tomoceridae 
Plutomurus californicus 
Troglophile 
Tomoceridae 
Pogonognathellus bidentatus 
Troglophile 
Diplopoda 
Chordeumatida 
Conotylidae 
Plumatyla sp. 
Troglobite 
0.50 
Striariidae 
Amplaria shastae 
Troglobite 
0.50 
Julida 
Paeromopodidae 
Californiulus dorsovittatus 
Epigean 
Parajulidae 
Bollmaniulus sp. 
Undetermined 
Undetermined 
Undetermined Diplopoda 
Troglobite 
0.04 
Diplura 
Dicellurata 
Japygidae 
Occasjapyx kofoidi 
Troglobite 
0.50 
Insecta 
Coleoptera 
Staphylinidae 
Aleocharinae 
Staphylinidae 
Sepedophilus castanea 
Epigean 
Tenebrionidae 
Eleodes productus 
Epigean 
Diptera 
Tipulidae 
Dactylolabis postiana 
Trogloxene 
Tipulidae 
Dicranoptycha sp. 
Trogloxene 
Tipulidae 
Tipula grahamina 
Trogloxene 
Lepidoptera 
Geometridae 
Triphosa haesitata 
Trogloxene 
Orthoptera 
Rhaphidophoridae 
Pristoceuthophilus n. sp. no. 4 
Troglophile 
Psocoptera 
Psyllipsocidae 
Psyllipsocus ramburii 
Troglophile 
Malacostraca 
Isopoda 
Ligiidae 
Ligidium kofoidi 
Troglobite 
0.07 
Chordata 
Amphibia 
Caudata 
Plethodontidae 
Hydromantes shastae 
Trogloxene 
Mammalia 
Carnivora 
Felidae 
Puma concolor 
Chiroptera 
Vespertilionidae 
Corynorhinus townsendii townsendii 
Trogloxene 
Rodentia 
Erethizontidae 
Erethizon dorsatum 
Trogloxene 
Mollusca 
Gastropoda 
Stylommatophora 
Bradybaenidae 
Monadenia churchi 
Epigean? 
Bradybaenidae 
Monadenia mariposa 
Epigean? 
Bradybaenidae 
Monadenia troglodytes 
Troglophile 
Haplotrematidae 
Haplotrema keepi 
Epigean? 
Helminthoglyptidae 
Helminthoglypta cypreophila 
Troglophile 
Pristilomidae 
Pristiloma cavator 
Troglophile 
Pristilomidae 
Pristiloma sp. 
Epigean? 
SE Score 5.92 
