ELLIOTT ET AL.: THE CAVE FAUNA OF CALIFORNIA 
151 
Family Planariidae 
Phagocata crenophila Carpenter (stygophile?) 
Records: Siskiyou Co.: Skunk Hollow Cave?; Trail Junction Cave. 
Comment: This species was originally described from springs in California, Nevada, and Utah. The cave population is a 
lighter color than the usually darkly pigmented spring populations (R. Kenk, pers. comm.). 
Bibliography: Carpenter (1969). 
Polycelis sp. (stygobite?) 
Records: Tulare Co.: Cirque Cave; Crystal Sequoia Cave; Panorama/Sink Cave System; White Chief Cave. 
Bibliography: Despain (2006); Krejca (2006, 2009b); Sluys (2009). 
PHYLUM NEMATODA (roundworms) 
Undetermined 
Records: Fresno Co.: Church Cave; Millerton Lake Cave System. Siskiyou Co.: Bigfoot Cave; Frozen Falls Cave. Tulare 
Co.: Hurricane Crawl Cave. 
Bibhography: Despain (1993); Graening (2011c). 
PHYLUM NEMATOMORPHA (Gordian or horsehair worms) 
Undetermined (parasite) 
Records: Tulare Co.: Lilbum Cave. 
Comment: In 2004 a horsehair worm was photographed by Jean Krejca in Lilbum Cave near water. The parasite probably 
erupted from a cave cricket (Figure x). 
PHYLUM ANNELIDA 
CLASS CLITELLATA (segmented worms) 
SUBCLASS OLIGOCHAETA 
ORDER HAPLOTAXIDA (earthworms) 
Undetermined 
Records: Calaveras Co.: *Cave City Cave. Fresno Co.: Boyden Cave; Church Cave; Millerton Lake Cave System. Santa 
Cruz Co.: Vanished River Cave; Dolloff Cave; IXL Cave; Vanished River Cave. Shasta Co.: *Potter Creek Cave. Siskiy¬ 
ou Co.: *Lyon’s Road Cave; Sugar Pine Butte Ice Cave. Tulare Co.: Crystal Sequoia Cave; Eighteenth Hole; Harry’s Bend 
Cave; Hurricane Crawl Cave; Kaweah Monkeyflower Cave; Lange Cave; Lilbum Cave; Overhang Cave; Panorama Cave; 
Paradise Cave; Seldom Seen Cave; Ursa Minor Cave; White Chief Cave. 
Comment: The earthworms and microdrile oligochaete families have been poorly documented, and deserve further study. 
Most of the records above were reported as Lumbricidae or Lumbricus. 
Bibhography: Bowers (2008a); Danehy (1951); Krejca (2006, 2009c); Lange (1954); Sinclair (1904); Taylor and Krejca 
(2006); Ubick (2001); Weaver (2008). 
ORDER LUMBRICULIDA 
Family Lumbriculidae (aquatic worms) 
Stylodrilus californianus Rodriguez (phreatophile) 
Record: Kern Co.: well RLSA 2204W (type-locality). 
Bibhography: Rodriguez (1996). 
ORDER TUBIFICIDA 
Family Naididae 
Comment: These undescribed species recently collected in Empire Cave appear to belong to the following subfamilies, but 
they are not clearly attributable to existing genera. Phallodrilines have not yet been reported in Nearctic caves, although 
many stygophilic species and genera have been described in Europe. (S.V. Fend, pers. comm.) 
Phallodrilinae n. sp. 
Record: Santa Cruz Co.: Empire Cave. 
Rhyacodrilinae n. sp. 
Record: Santa Cruz Co.: Empire Cave. 
