200 
PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 64, Supplement 1 
Catacombs Cave; Corkscrew Cave; Echoplex Cave; Frozen Falls Cave; Harris Mountain Cave; Fyon’s Road Cave; Maze 
Cave; Merrill Ice Cave; Mushpot Cave; Nirvana Cave; Santa Claus Cave; Post Office Cave; Red Tape Cave; Rollercoaster 
Cave; Spider Cave; Sugar Pine Butte Ice Cave; Tichnor Cave/Bertha’s Cupboard Cave; Township Cave; Trail Junction 
Cave; Upper Heppe Cave; Upstairs-Downstairs Cave. Trinity Co.: Paul Gibson Cave. Tulare Co.: Hidden Cave; Ursa 
Minor Cave. Tuolumne Co.: Crystal Palace. 
Comment: Some of these were badly damaged specimens. Specimens from Siskiyou County were taken from the middens 
and droppings of Neotoma, from the surface of small drip-pools, and under rocks. 
Bibliography: Krejca (2007); Taylor and Krejca (2006); Zara Environmental (2009). 
Foveacheles (Foveacheles ) auricularia Zacharda and Elliott (troglophile) 
Records: Calaveras Co.: Brown Deer Cave; Grapevine Gulch Cave. Siskiyou Co.: Planetary Dairy Cave; Trail Junction 
Cave (type locality). 
Comment: This species seems closely related to F. (F. ) osloensis (Sig Thor), a European troglophile. Zacharda (1980) and 
Zacharda and Elliott (1981, 1985) have discussed the zoogeography of post-glacial relicts of the family Rhagidiidae. Sev¬ 
eral species are epigean forms in the Arctic but have cave-limited populations in the temperate zone. It would not be sur¬ 
prising to find this species as an epigean form in northern Canada or Alaska. The air temperature in both of the Siskiyou 
County alpine caves was 3° C in late April, 1979, when the specimens were collected. 
Bibliography: Graening (2011c); Palacios-Vargas, Decu, Iavorski, Hutzu, and Juberthie (1998); Zacharda (1980); Zachar¬ 
da and Elliott (1981a, 1985). 
Foveacheles ( Usitorhagidia ) titanica Zacharda and Elliott (troglobite) 
Records: Calaveras Co.: Eagle View Cave No. 2. Tuolumne Co.: Crystal Palace; Indian Quarry Cave No. 1 (type local¬ 
ity). 
Comment: This is one of the largest rhagidiid species in the world (up to 2330 pm), and it bears a distinct troglobitic mor¬ 
phology. The chelicerae are long (up to 430 pm) and slender. Rhagidiids are adapted for feeding on collembolans and gen¬ 
erally have large chelicerae (Zacharda, 1980). This species is distributed on both sides of the Stanislaus River, as are the 
harvestmen Banksula grahami and B. melones. The highly insular distribution of cavemiferous limestone in this area prob¬ 
ably precludes dispersal of troglobites under the river canyon. Since the river probably began downcutting in its present 
position in the late Pliocene or early Pleistocene it seems likely that these populations have been isolated from one anoth¬ 
er since that time (Zacharda and Elliott, 1985). 
Bibliography: Culver, Christman, Elliott, Hobbs, and Reddell (2000); Palacios-Vargas, Decu, Iavorski, Hutzu, and 
Juberthie (1998); Zacharda and Elliott (1981, 1985). 
Poecilophysis ( Dentocheles ) melanoseta Zacharda and Elliott (troglophile) 
Records: Calaveras Co.: Diane’s Cave; Dirty Fissure; Poison Oak Cave. Mariposa Co.: Damp Cave. Riverside Co.: 
Mountaineer Mine. Tuolumne Co.: Indian Quarry Cave No. 1; Transplant Mine tailings; Vulture Cave (type locality). 
Comment: This species has also been found under stones in the Stanislaus River Canyon near the Indian Quarry Caves. 
Elliott (1978) and McEachem and Grady (1978) reported this as Rhagidia nr. cavernarum (Packard); however, this species 
is close to P. (D.) pratensis (C.L. Koch), which occurs in Europe, Alaska, Hawaii, and Japan (Zacharda and Elliott, 1981). 
Bibliography: Elliott (1978); McEachem and Grady (1978); Palacios-Vargas, Decu, Iavorski, Hutzu, and Juberthie (1998); 
Zacharda and Elliott (1981, 1985). 
Family Trombiculidae (chiggers) 
Undetermined 
Records: Calaveras Co.: Brown Deer Cave; Grapevine Gulch Cave. 
Comment: Immatures of this family are usually parasitic on vertebrates while the adults are free-living. 
Bibliography: McEachem and Grady (1978). 
Euschoengastia sp. (parasite) 
Records: Calaveras Co.: Cave City Cave. Mendocino Co.: Negro Hole Cave. Tulare Co.: Lost Soldier’s Cave. 
Tuolumne Co.: Crystal Palace. 
Tecomatlana (Tecomatlana ) watkinsi Vercammen-Grandjean (parasite) 
Records: Calaveras Co.: Cave of Skulls. Mariposa Co.: Bower Cave; Pool Pit. Riverside Co.: Alice Mine; Mountaineer 
Mine. Santa Cruz Co.: Empire Cave. Siskiyou Co.: Skunk Hollow Cave; Trail Junction Cave. Tulare Co.: Lost Soldier’s 
Cave. Tuolumne Co.: McLean’s Cave; Pinnacle Point Cave. 
