PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement I, pp. 1-279, 40 figs., 3 tables September 28, 2018 
United States Bat Species of Concern: A Synthesis 
Thomas J. O’Shea Paul M. Cryan 2 , Michael A. Bogan 3 
1 Scientist Emeritus, U.S. Geological Survey, Fort Collins Science Center, 2150 Centre Ave. Building C, 
Fort Collins, Colorado, 80526-8118 (Email: osheat@usgs.gov ); 2 Research Biologist, U.S. Geological 
Survey, Fort Collins Science Center (Email: cryanp@usgs.gov ); 3 U.S. Geological Survey (Retired), 
PO Box 2452, Corrales, NM 87048 (Email: mbogan@unm.edu ) 
Contents 
Abstract.2 
Introduction and Objectives.2 
Species of Concern.3 
Changes in the Federal Status of Bats in the United States and Territories.4 
A Need for Conservation and Research for U.S. Bats.5 
Objectives, Organization, and Content of this Publication.6 
Species Accounts: The Territories.9 
Pteropus samoensis — Samoan Flying Fox (Family Pteropodidae).9 
Stenoderma rufum — Red Fruit Bat (Family Phyllostomidae).14 
Species Accounts: The United States.17 
Choeronycteris mexicana — Mexican Long-tongued Bat (Family Phyllostomidae).17 
Corynorhinus rafinesquii — Rafmesque’s Big-eared Bat (Family Vespertilionidae).21 
Corynorhinus townsendii — Townsend’s Big-eared Bat (Family Vespertilionidae).37 
Euderma maculatum — Spotted Bat (Family Vespertilionidae).57 
Eumops perotis californicus — Greater Bonneted Bat (Family Molossidae).66 
Eumops underwoodi — Underwood’s Bonneted Bat (Family Molossidae).73 
Idionycteris phyllotis —Allen’s Big-eared Bat (Family Vespertilionidae).76 
Macrotus californicus — California Leaf-nosed Bat (Family Phyllostomidae).82 
Myotis austroriparius — Southeastern Myotis (Family Vespertilionidae).90 
Myotis ciliolabrum — Western Small-footed Myotis (Family Vespertilionidae).101 
Myotis evotis — Long-eared Myotis (Family Vespertilionidae).113 
Myotis leibii — Eastern Small-footed Myotis (Family Vespertilionidae).125 
Myotis occultus —Arizona Myotis (Family Vespertilionidae).136 
Myotis thysanodes — Fringed Myotis (Family Vespertilionidae).146 
Myotis velifer — Cave Myotis (Family Vespertilionidae).158 
Myotis volans — Long-legged Myotis (Family Vespertilionidae).169 
Myotis yumanensis — Yuma Myotis (Family Vespertilionidae).183 
Nyctinomops macrotis — Big Free-tailed Bat (Family Molossidae).196 
Summary and Highlights.203 
Acknowledgments.217 
Literature Cited.217 
Index.267 
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