O’SHEA, CRYAN & BOGAN: UNITED STATES BAT SPECIES OF CONCERN 
107 
et al., 2011a). In an earlier study at Mesa Verde National Park during 1989-1994, they also were 
intermediate in abundance, ranking fourth with 13 captures among 189 bats of 11 species (Chung- 
MacCoubrey and Bogan, 2003). This species was also intermediate in abundance (102 captured 
among 1,398 bats of 10 species, ranking sixth most frequently captured) among those taken in pon- 
derosa pine and Douglas fir forests along the Colorado Front Range, primarily in Boulder County 
(Adams et al., 2003), and seventh of nine species (14 bats among 634 individuals) in similar habi¬ 
tats in adjacent Larimer County (O’Shea et al., 2011b). In contrast, they were apparently absent 
from Engelmann spruce-subalpine fir forests at 2,900 meters in the central Rocky Mountains of 
Colorado (Storz and Williams, 1996) and were rarely captured in urbanizing areas at Fort Collins, 
Colorado (ranking fifth of seven species, with two bats among 504 individuals; O’Shea et al., 
2011b). In western Colorado, this species ranked seventh in abundance of 16 species (24 taken 
among 899 bats) captured at Colorado National Monument and the adjacent Mclnnis Canyons 
National Conservation Area during netting over small ephemeral pools in deep slickrock canyons 
within primarily pinon-juniper woodland and riparian habitats (Neubaum, 2017). Western small¬ 
footed myotis also ranked seventh in abundance (81 captures among 1,377 bats of 15 species) in 
mist-netting surveys at Dinosaur National Monument in northwestern Colorado and adjacent parts 
of Utah, at elevations ranging from 1,459 to 2,263 meters (Bogan and Mollhagen, 2016). 
Utah : Western small-footed myotis ranked thirteenth in relative abundance of 15 species 
(eight individuals out of 572 bats) in the Henry Mountains of Utah, where they were netted over 
water at 2,335 to 2,713 meters elevation (Mollhagen and Bogan, 1997). None were captured at 
Arch Canyon on the Colorado Plateau in southeastern Utah where 295 bats of 15 species were 
taken at elevations ranging from 1,474 to 1,707 meters (Mollhagen and Bogan, 2016). 
Wyoming: Western small-footed myotis ranked third among 12 species (43 captured out of 
about 370 individuals) documented by mist netting in lower elevation basin and foothills habitat in 
the south-central part of Wyoming during 2012 (Abemethy et al., 2013). They were the least abun¬ 
dant (one of 112 individuals of seven species) captured in late summer-early autumn 2010-2011 
by mist netting over water at elevations ranging from 1,568 to 3,116 meters in lodgepole pine, 
Engelmann spruce, subalpine fir, and Rocky Mountain juniper forests with open sagebrush and 
grassland habitats on the northern range of Yellowstone National Park, northwestern Wyoming 
(Johnson et al., 2017). They were not documented among 246 bats of six species captured in mist 
net surveys over streams and beaver ponds in and near the Medicine Bow National Forest in south¬ 
ern Wyoming, at elevations ranging from 2,133 to 2,896 meters and in habitats encompassing 
lodgepole pine ( Pinus contorta) and spruce-fir forests (Graver, 2002). 
South Dakota : Western small-footed myotis were the least abundant species captured during 
warm seasons in ponderosa pine-dominated habitat in the southern Black Hills of South Dakota, 
with 63 bats taken among 1,197 individuals of seven species (Cryan et al., 2000). However, they 
were the most common species in sampling at Badlands National Park in South Dakota (198 west¬ 
ern small-footed myotis out of 405 bats of nine species; Bogan et al., 1996, see also Famey and 
Jones, 1980). 
Alberta, Canada: Western small-footed myotis ranked as least abundant (two captures among 
1,868 individuals) of eight species of bats mist-netted over water in riparian habitats through urban 
Calgary and surrounding prairies in Alberta, Canada (Coleman and Barclay, 2012). However, 
Lausen and Schowalter (2008) provided a composite tabulation of results of mist netting in areas 
with suitable roosting habitat across Alberta, allowing a crude estimate that about 30% of 3,137 
bats captured in unpublished surveys were this species (but see notations on biases in Lausen and 
Schowalter, 2008). 
Elevational Differences in Habitats among Sex and Age Classes: Male and non-reproduc- 
