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PROCEEDINGS OF THE CALIFORNIA ACADEMY OF SCIENCES 
Series 4, Volume 65, Supplement I 
species (Storz and Williams, 1996). In western Colorado, this species ranked fourteenth in relative 
abundance of 16 species (two among 899 bats) captured at Colorado National Monument and the 
adjacent Mclnnis Canyons National Conservation Area during netting over small ephemeral pools 
in deep slickrock canyons within primarily pinon-juniper woodland and riparian habitats 
(Neubaum, 2017). 
In Boulder County, Colorado, long-eared myotis were moderately abundant in ponderosa pine 
and Douglas fir/mixed conifer forests, ranking third in abundance among 10 species and 1,398 
individuals taken at the Boulder County sites (Adams et al., 2003). They ranked sixth of nine 
species (38 bats among 634 individuals) in similar habitats in adjacent Larimer County, Colorado 
but were most abundant above 2,000 meters (O’Shea et al., 2011b). This species ranked third in 
abundance (162 captures among 1,377 bats of 15 species) in surveys at Dinosaur National Monu¬ 
ment in northwestern Colorado and adjacent parts of Utah at elevations ranging from 1,459 to 
2,263 meters (Bogan and Mollhagen, 2016). 
Utah: Long-eared myotis ranked second in relative abundance of 15 species (75 captures 
among 572 individuals) in the Henry Mountains of southeastern Utah, where elevations of capture 
sites where this species was taken ranged from 1,433 to 2,713 meters (Mollhagen and Bogan, 
1997). In contrast, at Arch Canyon on the Colorado Plateau in southeastern Utah these bats were 
among the least abundant species, with two bats captured among 295 bats of 15 species taken at 
elevations ranging from 1,474 to 1,707 meters (Mollhagen and Bogan, 2016). 
Wyoming: Long-eared myotis ranked second of seven species (23 of 112 individuals) captured 
in late summer-early autumn 2010-2011 by mist netting over water at elevations ranging from 
1,568 to 3,116 meters in lodgepole pine, Engelmann spruce, subalpine fir, and Rocky Mountain 
juniper forests with open sagebrush and grassland habitats on the northern range in Yellowstone 
National Park, Wyoming (Johnson et al., 2017). During 2012 they ranked highest among 12 species 
(162 captured among about 370 individuals) documented by mist netting in lower elevation basin 
and foothills habitat in the south-central part of Wyoming (Abemethy et al., 2013). They were low 
in relative abundance (five among 246 bats of six species, ranking fifth) of bats captured in mist 
net surveys over streams and beaver ponds in and near the Medicine Bow National Forest in south¬ 
ern Wyoming, at elevations ranging from 2,133 to 2,896 meters and in habitats encompassing 
lodgepole pine (Pinus contorta) and spruce-fir forests (Graver, 2002). 
Elevational Differences in Habitats among Sex and Age Classes: A higher proportion of 
males were found at elevations greater than 2,311 meters (65%) than below 2,165 meters (15%) in 
a sample of 270 long-eared myotis from Mesa Verde National Park in southwestern Colorado 
(O’Shea et al., 2011a). Sex ratios were equal in a sample of 218 adults captured at elevations rang¬ 
ing from 1,350 to 2,150 meters in predominantly lodgepole pine forests in and around the 
Kananaskis Valley, Alberta, Canada (Barclay, 1991). 
Foraging and Dietary Analysis. — In red fir (Abies magnifica ) - lodgepole pine forests in 
the Sierra Nevada of California, foraging individuals have been described as flying in straight 
courses in open spaces about 12 meters above ground during early evening, hunting closer to the 
ground later in the evening (Ingles, 1949). In coniferous forests (predominantly lodgepole pine) of 
the Rocky Mountains in Alberta, long-eared myotis foraged mostly along paths through or within 
the forest (Barclay, 1991). Activity areas of foraging M. evotis radio tracked in the western Cas¬ 
cades of Oregon averaged 38 ha, were a mean distance of 518 meters from the day roosts, and were 
significantly closer to water than random points (Waldien and Hayes, 2001). 
Reproductive females studied among cottonwood groves along the Saskatchewan River Val¬ 
ley in Alberta foraged nearly all night long every night with little night roosting, regardless of sea¬ 
sonal differences in the length of darkness, suggestive of narrow energy budgets (Chraszcz and 
