DANIEL & TRIPP: LOUTERIDIUM : TAXONOMY, BIOLOGY, AND CONSERVATION 77 
image seen); Canton Siquirres, La Alegria, Alto Botella, [ca. 10°05’45.64”N, 083°36’58.36”W], 
J. Gonzalez et al. 7967 (LSCR-image seen), 9618 (LSCR-image seen); Canton Siquirres, Florida, 
Alto Botella, [10°05’19.97”N, 083°33 , 26.45”W], J. Gonzalez et al. 9353 (LSCR-image seen); 
Canton Siquirres, Pocora, Cuenca del Rio Destierro, Cordillera de Talamanca, Canton de Matina, 
200 m aguas abaja de la confluencia de Quebrada Caiiabral con Rio Barbilla, 10°00’10”N, 
083°25’30”W, G. Herrera 2286 (DUKE, MO); Almirante, Cerro Chiqui, 09°43’40”N, 
083°18’30”W, G. Herrera & W. Gamboa E. 8581 (F, K, MEXU, MO, NY, US); 10 km SW of 
Siquirres on road to Turrialba, E of ridge top, J. MacDougal 1110 (DUKE); Canton de Limon, 
Cordillera de Talamanca, Cerro Muchilla, Fila Matama, entrando por pueblo El Progresso, 
09°47’40”N, 083°06’30”W, R. Robles &A. Chacon 2765 (MO). Puntarenas: vicinity of Finca Las 
Cruces, SE of San Vito on road to ViaNeilly, [ca. 08°47’14.82”N, 082°57’25.04”W], R. Weaver et 
al. 1777 (DUKE). San Jose: from La Montura to Los Chorritos, L. Gomez et al. 20929 (MO). 
PANAMA. Bocas del Torro: Fish Creek Mts., vicinity of Chiriqui Lagoon [ca. 08 o 58’59.68”N, 
082°13’48.70”W], H. von Wedel 2283 (MO). Chiriqui: Fortuna Dam site [ca. 08°44’46.90”N, 
082°14’38.50”W], J. Folsom & R. Dressier 5464 (MO); La Fortuna hydroelectric project, ridge 
behind camp, [OS^l^.O^’N, 082°13’47.82”W], B. Hammel 2228 (MEXU, MO). Colon: along 
Rio Guanche, 3-7 km above bridge [ca. 09°29’N, 079°38’W], B. Hammel et al. 4916 (MO); ca. 2- 
3 mi up Rio Guanche [ca. 09°30’30”N, 079°39’30”W], H. Kennedy & R. Foster 2180 (MO). Guna 
Yala (San Bias): Nusagandi, El Llano-Carti Road, along creek on Atlantic slope [ca. 
09°18’18.56”N, 078°59’20.14”W], G. de Nevers & B. Gonzalez 3656 (DUKE, MEXU, MO); 
El Llano-Carti Road, KM 19.1 [ca. 09 o 20’51.93”N, 079°01 , 54.94”W], G. de Nevers et al. 6195 
(CAS, DUKE, MEXU, MO); trail to Cerro Obu (Habu of maps) from Rio Urgandi (Rio Sidra) 
[ca. 09°23’59.51”N, 078°49’40.88”W], G. de Nevers et al. 8007 (CAS, MO); El Llano-Carti hwy., 
ca. 23 km N of El Llano, R. Dressier 4310 (DUKE, ENCB, F, MO); Nusagandi, El Llano-Carti 
road, 09°19’N, 078°55’W, H. van der Werff7019 (CAS, MO). 
4. Louteridium dendropilosum T.F. Daniel, Proc. Calif. Acad. Sci. 64:140. 2017. Type.— 
MEXICO. Oaxaca: Distr. Pochutla, Mpio. San Miguel del Puerto, Arroyo Arena, ca. 100 m down¬ 
stream from jet. Rio Laja, ca. 3 km SE of Rancho Dioon toward Xadani, 15°58’51.33”N, 
096°05’53.9r’W, 600 m, evergreen seasonal forest (selva mediana subperennifolia), 29-III-2011, 
T. Daniel, A. Sanchez, andJ. Pascual 11784 (holotype: MEXU!; isotypes: CAS! COLO! K! MO! 
NY! SERO! US!). 
Shrubs to trees to 12 m tall, sometimes epipetric. Older (woody) stems quadrate, lenticellate, 
irregularly striate-sulcate, lacking trichomes; younger (herbaceous) stems subquadrate-sulcate, 
sparsely lenticellate, irregularly fissured, evenly pubescent with erect to flexuose simple and den¬ 
dritic (sparse) eglandular trichomes <0.1-0.5 mm long. Leaves seasonally deciduous, ± clustered 
at apex of old growth or at apex of an otherwise leafless shoot of new growth, petiolate, petioles to 
65 mm long, blades subsucculent, ovate to elliptic to broadly elliptic, 76-190 mm long, 40-122 
mm wide, 1.4-2.5 x longer than wide, apiculate to acuminate at apex, rounded to acute to attenu¬ 
ate at base, adaxial surface pubescent with simple and dendritic eglandular trichomes, trichomes 
soon becoming ± restricted to proximal portion or to midvein, abaxial surface pubescent (especially 
along veins) with dendritic trichomes to 0.5 mm long, midvein often pinkish or reddish, margin 
entire (sometimes undulate and appearing subcrenate). Inflorescence a terminal subsessile to 
pedunculate raceme to 220 mm long, peduncle to 50 mm long, pubescent like young stems, rachis 
pubescent like young stems; dichasia opposite or alternate, sessile, 1-flowered, to 47 mm long 
(excluding corollas). Bracts caducous, lanceolate to lance-ovate, 11-16 mm long, 2.5-6 mm wide, 
abaxially pubescent with simple and dendritic trichomes 0.05-0.2 mm long. Bracteoles caducous, 
