DANIEL & TRIPP: LOUTERIDIUM : TAXONOMY, BIOLOGY, AND CONSERVATION 85 
J. Morales & M. Garda 4377 (MO); Cerro Chiblac, between Finca San Rafael and Ixcan, Sierra 
de los Cuchumatanes [ca. 15°46’25”N, 091°13 , 58”W], J. Steyermark 49180 (F, US). Izabal: 
25 km SSW of Puerto Barrios, [ca. 15°34’21.26”N, 088°37 , 0.05”W], D. Gregory 612 (CAS, F, 
US); Modesto Mendez, [ca. 15 0 53’29.49”N, 089°13’46.15”W], W Harmon & J. Fuentes 2089 
(MO); Biotopo Chocon Machacas, Livingston, 15°45’56”N, 088°55’55”W, J. Morales 1959 (MO); 
Rio Dulce, 2-A mi W of Livingston, [ca, 15°48’12.69 ,, N, 088 o 46’37.09”W], J. Steyermark 39562 
(F); Chocon River, S. Watson 292 (US). Peten: Dolores, S of village, [ca. 16°30 , 51.17”N, 
089°24’56.87”W], E. Contreras 2097 (LL, MO, US); N of Lake Macanche, foot of Cerro Rocoso 
at Mancanche [ca. 16°58’04”N 089°38’20”W], E. Contreras 5510 (BM, F, K, MO, NY, WIS, US); 
La Cumbre, W of KM 145, [ca. 16°04’51.97”N, 089°21’3.37”W], E. Contreras 6646 (F, LL, MO, 
PH, US); Tikal District, ruins, Naranjo, O. Cook & R. Martin 79 (US); along Rio Dulce-Flores 
highway near KM 327, ca. 10 km NW of San Pedro Cadenas, 15°58’08.6”N, 089 o 16’42.7”W, 
T. Daniel & M. Veliz 11255 (BIGU, CAS, MICH, US); Strasse von Flores zur Rio Dulce-Bracke, 
Strecke zwischen Poptun und San Luis, 12 km S von Poptun, H. Kurz & R. Frisch 147 (W), 148 
(W); Sayab, Fallabon-Yaxha road, [ca. 17°04’01.49”N, 089°22’56.17”W], C. Lundell 2081 
(F, MICH); Arroyo Dolores, ca. 200 m de Dolores, R. Ortiz 1569 (F, MO, NY, US); San Luis, [ca. 
16°11’37.30”N, 089°25’48.59”W], P. Ventura 177 (F). Zacapa: Volcan de Monos, [ca. 
15°08’58.95”N, 089°30’18.66”W], J. Steyermark 42400 (F, US). Department undetermined: east¬ 
ern portions of Vera Paz and Chiquimula, S. Watson s.n. (K). HONDURAS. Comayagua: Villa de 
Taulabe, Quebrada la Caliche, [ca. 14°39’56.69”N, 087 o 59’26.20”W], C. Alduvin 101 (MO); 
Siguatepeque, [ca. 14°36’ 10.86”N, 087°47’54.39”W], J. Edwards P-589 (F); Quebrada el Caliche, 
SE de la Villa Taulabe, [ca. 14°41’02.38”N, 087°57’02.73”W], D. Ruiz 138 (NY, TEFH); near 
Taulabe, P. Standley et al. 7001 (F); Pito Solo, Lago Yojoa, [ca. 14°47’39.06”N, 087°58’38.47”W], 
J. Valerio R. 2940 (F); near Pito Solo, L. Williams & A. Molina R. 12335 (F); along Lake Yojoa, 
near Pito Solo, L. Williams & A. Molina R. 17730 (F). Copan: Montana Los Zapotes, W of Los 
Zapotes, 18 km N of Copan Ruinas on road to Agua Caliente and 2-3 km W of Los Zapotes school, 
[ca. 14°59’13.83”N, 089°09’04.57”W], T. Hawkins 981 (CAS, MO); Montana Espiritu Santo, 
15°05’N, 088°55’W, T. Hawkins & D. Mejia 185 (EAP, MO). Cortes: entre Pito Solo y Agua Azul, 
Lago Yojoa, [ca. 14°51’07.26”N, 087°57’14.09”W], A. Molina R. 10619 (EAP, F); camino entre 
Tapiquilares y Las Crucitas, 20 km S de San Antonio de Cortes, al pie de la montana La Nieve, [ca. 
14°59’22.86”N, 088°02’19.81”W], C. Nelson et al. 8128 (MO). Santa Barbara: eastern slopes of 
Cerro Santa Barbara, [ca. 14 0 53’14.92”N, 088°07’01.01”W], P. Allen et al. 6047 (CAS, EAP, F, 
GH, US); Cerro Sta. Barbara, R. Armour & A. Chable 6047 (BM, F); Punta Gorda, Lago Yojoa, 
14°53’N, 088°00’W, R. Evans 1001 (CAS); Punta Gorda, Lago Yojoa, 14°53’N, 088°00’W, 
R. Liesner 26754 (MO). 
6. Louteridium koelzii Miranda & McVaugh, Ann. Inst. Biol. Mex. 32:182. 1962 (“1961”). 
Type.— MEXICO. Jalisco: 8 mi SW of Pihuamo [ca. 19°11’26.09”N, 103°28’3.97”W], 500-600 
m, 6-XII-1959 (fir), R. McVaugh & W. Koelz 1507 (holotype: MICH!; isotype: MEXU!). 
Trees to 10 m tall. Older (woody) stems subquadrate, irregularly fissured-striate; younger 
(herbaceous) stems stubby, quadrate, glabrous. Leaves deciduous, clustered at apex of old growth, 
petiolate, petioles to 73 mm long, blades membranaceous, ovate to elliptic, 90-220 mm long, 
35-100 mm wide, 2-2.7 x longer than wide, acute to acuminate at apex, rounded to attenuate and 
asymmetric at base, surfaces glabrous, margin entire. Inflorescence of axillary and/or terminal 
subsessile to pedunculate thyrses (main axis with lateral pedunculate dichasia appearing as branch¬ 
es) to 250 mm long, peduncle to 50 mm long, pubescent with erect to flexuose eglandular 
